Calliophis / Hemibungarus - Oriental coral snakes
Ota et al. (1999) investigated patterns
of variation in the Hemibungarus japonicus complex in the Ryukyu
Islands and on Taiwan. Three subspecies of H. japonicus are recognised
from the Ryukyus: H.j. japonicus from Amamioshima and adjacent islands;
H.j.
boettgeri from Tokunoshima, Okinawa and adjacent islets; and
H.j.
takarai from Kunejima and several islets north of Okinawa. On Taiwan,
two separate species were recognised: H. hatori from the northern
half of Taiwan, and H. sauteri from the southern half of the island.
The generic classification of Asiatic
coral snakes requires further research. All were formerly included in the
genus Calliophis (e.g., Smith, 1943; Golay, 1985). However, based
on differences in jaw musculature, McDowell (1987) informally suggested
that the Oriental coral snakes should be split into three genera, and this
was implemented by Golay et al. (1993) and David & Ineich (1999): the
genus Calliophis contains the species C. bibroni, C. gracilis,
and C. melanurus; Hemibungarus contains H. calligaster,
H. hatori, H. japonicus, H kelloggi, H. macclellandi, and H. sauteri;
Finally, the species formerly known as Calliophis beddomei, C. maculiceps
and C. nigrescens were assigned to Maticora by Golay et al.
(1993). Further, more rigorous analyses of the phylogenetic relationships
among Asian coral snakes are needed to test this classification. See
later review by Slowinski et al (2001)
Elapsoidea trapei - African
garter snakes
Mane (1999) described Elapsoidea trapei from extreme southeastern Senegal. The new species differs from sympatric E. semiannulata moebiusi in having 6 rather than 7 supralabials, the 2nd and 3rd (rather than 3rd and 4th) in contact with the eye, and a dark brown rather than light venter, as well as more subtly in other chracateritsics.This brings the total number of Elapsoidea species known from African to 10: E. boulengeri (southern parts of Africa), E. broadleyi (Somalia), E. chelazziorum (Somalia), E. guentheri (south-central Africa), E. laticincta (Central Africa), E. loveridgei (Central and East Africa), E. nigra (Tanzania), E. semiannulata (widespread south of Sahara), E. sunderwallii (southern Africa) and E. trapei (Senegal).
Micrurus frontalis complex
- South American coral snakes
Micrurus frontalis
Jorge da Silva & Sites (1999) revised the classification of the Micrurus frontalis complex. Based on their analysis of various aspects of colour pattern and other morphological characters, they recognised the following seven species: Micrurus frontalis, from central Brazil (São Paulo, Minas Gerais, Mato Grosso, Mato Grosso do Sul, Goiás, Espírito Santo) and eastern Paraguay; M. altirostris from southern Brazil (Paraná, Santa Catarina, Rio Grande do Sul), eastern Paraguay, Uruguay and northwestern Argentina (Misiones, Corrientes and northern Entre Ríos Provinces); Micrurus baliocoryphus from northeastern Argentina and southwestern Paraguay; M. brasiliensis from northern central Brazil (northern Goiás, southern Tocantins and Bahia); Micrurus diana from Santa Cruz Province, Bolivia; M. pyrrhocryptus, from Argentina (south to Río Negro Province) as well as northwestern Paraguay and southern Bolivia; and Micrurus tricolor, from Brazil (Mato Grosso do Sul) and eastern Bolivia. Micrurus frontalis multicinctus is a synonym of M. altirostris.