Dr Matt Hayward - Staff Profile

Matt HaywardDr Matt Hayward
Senior Lecturer in Conservation

Room G2a, Thoday Building
Email: m.hayward@bangor.ac.uk


Tel.: 01248 383642 (from U.K.)
+44 1248 383642 (International)

Website: http://conservation.bangor.ac.uk/
Facebook: https://www.facebook.com/ConservationEcologyAtBangorUni
Twitter: @ConservResearch

Qualifications

Ph.D. (University of New South Wales, Australia) – 2003
B.Sc. Hons 1 (UNSW) - 1995

Research Interests

I am interested in the conservation ecology of threatened species, the factors that threaten them and the methods we can use to effectively conserve them. I have researched these conservation issues in Australia, South Africa and Poland on marsupials, rodents, reptiles, invertebrates, ungulates and large predators.  I have published on predator-prey interactions, reintroduction biology, population dynamics, spatial ecology, intra-guild competition, diet, ecosystem services, conservation effectiveness and status assessments.  I also have experience in conservation management (reintroduction, pest animal control, conservation fencing, fire management) and have sat on several Australian threatened species recovery teams.

Biography

I conducted a PhD on the conservation ecology of the vulnerable quokka – a small wallaby that the introduced red fox loves to kill – in the Western Australian jarrah forest. I then conducted two post docs in South Africa; the first on bushmeat hunting in the coastal forests of the Transkei with the Walter Sisulu University, and the second at the Nelson Mandela Metropolitan University to study the reintroduction of lions, spotted hyaenas and a leopard to Addo Elephant National Park.  After this I undertook a Marie Curie Post-Doctoral Fellowship at the Polish Academy of Science’s Mammal Research Institute in Białowieża Primeval Forest.  Most recently I have been the Australian Wildlife Conservancy’s regional ecologist for six reserves in south-eastern Australia covering over 700,000ha and ranging from the deserts of Lake Eyre through the mallee to Sydney’s North Head where reintroduction, ecosystem services, feral eradication/control and fire management were key research issues.

Editorial roles

Frontiers in Ecology and Evolution, Endangered Species Research, Wildlife Biology, Journal of Zoology, South African Journal of Wildlife Research, Journal of Applied Ecology, Conservation Letters

 

Offices held

Committees:

  • SBS Board of Studies
  • SENRGy Board of Studies
  • SENRGy Marketing Committee
  • CNS/CoPAS Marketing Committee

Roles:

  • Director of Zoology with Conservation degree programme
  • Director of Ecology degree programme
  • Director of 3rd Year Studies

Governance of Conservation/Charitable Organisations

  • Trustee of the Welsh Mountain Zoo (National Zoo of Wales)
  • Trustee of the Red Squirrels Trust Wales

Affiliations

  • Centre for African Conservation Ecology, Nelson Mandela University, South Africa (2006 – on-going)
    o School of Biological, Earth and Environmental Science, University of New South Wales, Australia (2003 – 2009)
    o Centre for Wildlife Management, University of Pretoria, South Africa (2009 – on-going)
    o School of Life Sciences, University of kwaZulu-Natal, South Africa (2014-2017)

Research Students

  • Ludmila Osipova – Corridor use by African elephants in East Africa. Co-supervised with Niko Balkenhol of Georg-August University, Goettingen, German.
  • Jeremy Ringma – Does competition with burrowing bettongs limit woylies: how and why? PhD co-supervised with Richard Fuller at University of Queensland, Australia.
  • Nicole Coggin – Impact of feral mammal eradication and native species reintroduction on the invertebrate fauna of Scotia Sanctuary.PhD co-supervised with Heloise Gibb of La Trobe University, Australia.
  • Peter Haswell – Are the impacts of apex predators context dependent: wolves in Croatia
  • Maarten Hofman – Corridor creation in Belize based on peccary spatial ecology. Co-supervised with Niko Balkenhol of Gottingen University, Germany
  • Carolyn Devens – Human-carnivore conflict in South Africa: how leopards persist in rangelands. Co-supervised with Michael Somers of the University of Pretoria.
  • Wafa’a Behbehani – Conservation ecology of the Arabian spiny-tailed lizard in Kuwait.
  • Tom Major – Conservation ecology of the adder in Wales. Co-supervised with Wolfgang Wüster of Bangor University.

Click here to visit the Research Student's profiles

Publications

Journal articles since 2005

2016

Hayward, M. W., W. J. Ripple, R. D. Plotz, & S. T. Garnett (2016). Don’t bank African rhino in Australia. Nature In press.

Creel, S., M. Becker, D. Christianson, E. Dröge, N. Hammerschlag, M. W. Hayward, U. Karanth, A. Loveridge, D. W. Macdonald, W. Matandiko, J. M’soka, D. Murray, E. Roseblatt & P. Schuette (2016) Policy for hunting large carnivores can and should be improved. Science, 350, eLetter: https://d2ufo47lrtsv5s.cloudfront.net/sites/default/files/response_attachments/2016/06/Creel_et_al.pdf

Ripple, W.J. G. Chapron, J. V. Lopez-Bao, S. M. Durant, D.W. Macdonald, P.A. Lindsey, E. L. Bennett, R. Beschta, J. T. Bruskotter, A. Campos-Arceiz, R. T. Corlett, C. T. Darimont, A. J. Dickman, R. Dirzo, H. T. Dublin, J. A. Estes, K. T. Everatt, M. Galetti, V. R. Goswami, M. W. Hayward, S. Hedges, M. Hoffmann, L. T. B. Hunter, G. I. H. Kerley, M. Letnic, T. Levi, F. Maisels, J. C. Morrison, M. P. Nelson, T. M. Newsome, L. Painter, R. M. Pringle, C. J. Sandom, J. Terborgh, A. Treves, B. Van Valkenburgh, J. A. Vucetich, A. J. Wirsing, A. D. Wallach, C. Wolf, R. Woodroffe, H. Young & L. Zhang (2016). Saving the world’s terrestrial megafauna. BioScience In press.

Linnell, J. D. C., A. Trouwborst, L. Boitani, P. Kaczensky, D. Huber, S. Reljic, J. Kusak, A. Majic, T. Skrbinsek, H. Potocnik, M. W. Hayward, E. J. Milner-Gulland, B. Buuveibaatar, K. A. Olson, L. Badamjav, R. Bischof, S. Zuther, U. Breitenmoser (2016) Border security fencing and wildlife: the end of the transboundary paradigm in Eurasia? PLoS Biology In press.

Moll, R., Killion, A. K., Montgomery, R. A., Tambling, C. J. & Hayward, M. W. (2016) Spatial patterns of African ungulate aggregation reveal complex but limited risk effects from reintroduced carnivores. Ecology 97, 1123-1134.

Luther, D. A., Brooks, T. M., Butchart, S., Hayward, M. W., Kester, M., Lamoreux, J. & Upgren, A. (2016) Determinants of bird conservation action implementation and associated population trends of threatened species. Conservation Biology In press.

Van Valkenburgh, B., Hayward, M. W., Ripple, W. J., Meloro, C. & Roth, V. L. (2016). The impact of large terrestrial carnivores on Pleistocene ecosystems. Proceedings of the National Academy of Sciences 133, 862-867. 
- F1000 Prime recommended: http://f1000.com/prime/725882781?bd=1&ui=21606

Hayward, M. W., Ward-Fear, G., L’Hotellier, F., Herman, K., Kabat, X. (2016). Could biodiversity loss have increased Australia’s bushfire threat? Animal Conservation In press.

Coggan, N. V., Hayward, M. W., & Gibb, H. (2016) Termite activity and decomposition are influenced by digging mammal reintroductions along an aridity gradient. Journal of Arid Environments, 133, 85-93

Eldred, T., Meloro, C., Scholtz, C., & Hayward, M. W. (2016). Does size matter for horny beetles? A geometric morphometric analysis of interspecific and intersexual size and shape variation in Colophon haughtoni Barnard, 1929, and C. kawaii Mizukami, 1997 (Coleoptera: Lucanidae). Organisms Diversity and Evolution, In press.

Hayward, M. W., Kamler, J. F., Montgomery, R. A., Newlove, A., Rostro-Garcia, S., Sales, L. P. & Van Valkenburgh, B. (2016) Prey preferences of the jaguar Panthera onca reflect the post-Pleistocene demise of large prey. Frontiers in Ecology and Evolution 3, e148 http://journal.frontiersin.org/article/10.3389/fevo.2015.00148/full

Haswell, P. M., Kusak, J. & Hayward, M. W. (2016). Large carnivore impacts are context-dependent. Food Webs In press.

Sales, L. P., Hayward, M. W., Zambaldi, L., Passamani, M., de Melo, F. R. , Loyola, R. (2016). Time-lags in primate occupancy: a study case using dynamic models. Natureza & Conservação, In press.

Gray, J., J. Brockington, M. W. Hayward & J. D. Walmsley (2016) How the proposed reintroduction of Eurasian lynx to Britain illustrates competing values and contrasting views associated with humans and the natural world. . Country-side, 36, In press.

2015

Creel, S., M. Becker, D. Christianson, E. Dröge, N. Hammerschlag, M. W. Hayward, U. Karanth, A. Loveridge, D. W. Macdonald, W. Matandiko, J. M’soka, D. Murray, E. Roseblatt & P. Schuette (2015) Questionable policy for large carnivore hunting. Science, 350, 1473-1475.

Ripple, W. J., Newsome, T. M., Wolf, C., Dirzo, R., Everatt, K. T., Galetti, M., Hayward, M. W., Kerley, G. I. H., Levi, T., Lindsey, P. A., Macdonald, D. W., Malhi, Y., Painter, L. E., Sandom, C. J., Terborgh, J. & Van Valkenburgh, B. (2015). Collapse of the world’s largest herbivores. Science Advances 1, e1400103 http://advances.sciencemag.org/content/1/4/e1400103

Eldridge, D.J., Woodhouse, J., Curlevski, N., Hayward, M. W., Brown, M. & Neilan, B. (2015). Soil foraging animals alter the composition and co-occurrence of microbial communities in a desert shrubland. ISME Journal 9, 2671-2681 http://www.nature.com/ismej/journal/vaop/ncurrent/full/ismej201570a.html

Hayward, M. W., Child, M. F., Kerley, G. I. H., Lindsey, P. A., Somers, M. J. , Burns, B. (2015). Ambiguity in guideline definitions introduces assessor bias and influences consistency in IUCN Red List status assessments. Frontiers in Ecology and Evolution 3, e87 http://dx.doi.org/10.3389/fevo.2015.00087.

Eldridge, D.J., Woodhouse, J., Curlevski, N., Hayward, M. W., Brown, M. & Neilan, B. (2015). Soil foraging animals alter the composition and co-occurrence of microbial communities in a desert shrubland. ISME Journal In press.

Ripple, W.J., Newsome, T.M., Wolf, C., Dirzo, R., Everatt, K.T., Galetti, M., Hayward, M.W., Kerley, G.I.H., Levi, T., Lindsey, P.A., Macdonald, D.W., Malhi, Y., Painter, L.E., Sandom, C.J., Terborgh, J. and Van Valkenburgh, B. (2015). Collapse of the world's largest herbivores. Science Advances 1, e1400103.

Silvey, C.J., Hayward, M.W. and Gibb, H. (2015). Effects of reconstruction of a pre-European vertebrate assemblage on ground-dwelling arachnids in arid Australia. Oecologia 178, 497-509.

2014

Clements, H.S., Tambling, C.J., Hayward, M.W. and Kerley, G.I.H. (2014). An objective approach to determining the weight ranges of prey preferred by and accessible to the five large African carnivores. PLoS ONE 9, e101054.

Durant, S., Pettorelli, N., Blackburn, T., Baillie, J., Boxshall, G., Brooks, S., Carbone, C., Crees, J., Cunningham, A., Daniels, M., Deinet, S., Ewen, J., Hayward, M., Macdonald, D. W., Mace, G., May, R., Norris, K., Pimm, S., Raffaelli, D. & Sutherland,W. (2014). UK bill could prompt biodiversity loss. Nature 512, 253.

Harris, S., Arnall, S., Byrne, M., Coates, D., Hayward, M. W., Martin, T., Mitchell, N. & Garnett, S. (2014). Whose backyard? Some precautions in choosing recipient sites for assisted colonisation of Australian plants and animals. Ecological Management and Restoration 14, 106-111.

Hayward, M.W., Lyngdoh, S. and Habib, B. (2014). Diet and prey preferences of dholes (Cuon alpinus): dietary competition within Asia's apex predator guild. Journal of Zoology 294, 255-266.

Hayward, M.W. and Marlow, N. (2014). Will dingos really conserve wildlife and can our methods tell? Journal of Applied Ecology 51, 835-838.

Lyngdoh, S., Shrotriya, S., Goyal, S. P., Clements, H., Hayward, M. W. & Habib, B. (2014). Prey preferences of the snow leopard (Panthera uncia): regional diet specificity holds global significance for conservation. PLoS ONE 9, e88349.

2013

Lunt I. D., Byrne M., Hellmann J. J., Mitchell N. J., Garnett S. T., Hayward M. W., Martin T. G., McDonald-Madden E., Williams S. E. & Zander K. K. (2013). Using assisted colonisation to conserve biodiversity and restore ecosystem function under climate change. Biological Conservation 157, 172-177.
Winterbach, H., Winterbach, C. W., Somers, M. J. & Hayward, M. W. (2013). Key principles related to the conservation management of large carnivores in Africa. Mammal Review 43, 89-110.
Jooste, E., Hayward, M. W., Pitman, R. & Swanepoel, L. (2013). Effect of prey mass and selection on predator carrying capacity estimates. European Journal of Wildlife Research 59, 487-494.

2012

Tambling, C. J., Druce, D. J., Hayward, M. W., Castley, J. G. & Kerley, G. I. H. (2012). Spatial and temporal changes in group dynamics and range use enable anti-predator responses in African buffalo. Ecology 93, 1297-1304.
Martin, T. G., Nally, S., Burbidge, A. A., Arnall, S., Garnett, S., Harris, S., Hayward, M. W., Holsworth, M., Lumsden, L. F., Menkhorst, P., McDonald-Madden, E. & Possingham, H. P. (2012). Acting fast avoids extinction: plight of the Christmas Island pipistrelle and Orange-bellied Parrot. Conservation Letters, 5, 274-280.
Hayward, M. W., Jędrzejewski, W. & Jędrzejewska, B. (2012). Prey preferences of the tiger Panthera tigris. Journal of Zoology 286, 221-231. 
Jedrzejewski, W., Niedzialkowska, M., Hayward, M. W., Goszczynski, J., Jedrzejewska, B., Borowik, T., Barton, K., Nowak, S., Harmuszkiewicz, J., Juszczk, A., Kalamarz, T., Kloch, A., Koniuch, J., Kotik, K., Myslajek, R. W., Nedzynska, M., Olczyk, A., Teleon, M. & Wojtulewicz, M. (2012) Prey choice and diet of wolves related to ungulate communities and wolf subpopulations in Poland. Journal of Mammalogy, 93, 1480-1492.
Hayward, M. W. & Hayward, M. D. (2012). Waterhole use by African fauna. South African Journal of Wildlife Research 42(2), 117-127.
Hayward, M. W., Kerley, G. I. H., Somers, M. J., Perrin, M., Bester, M. N., Dalerum, F., Do Linh San, E., Hoffman, L., Marshal, J. P., Mills, M. G.L, Nel, J. A. J. & Owen-Smith, N. (2012). Animal ethics and ecotourism. South African Journal of Wildlife Research 42(2), iii-iv.
Hayward, M. W. (2012). Time to agree on a conservation benchmark for Australia. Pacific Conservation Biology 18, 69-76.
Hayward, M. W., l’Hotellier, F., O’Connor, T., Ward-Fear, G., Cathcart, J., Cathcart, A., Stephens, A. J., Stephens, J., Herman, K. & Legge, S. (2012). Reintroducing bridled nailtail wallabies Onychogalea fraenata beyond conservation fences in New South Wales: Phase 1 of the reintroduction to Scotia Sanctuary – a male-only affair. Proceedings of the Linnean Society of New South Wales 134, A27-A37.
Eldridge D. J., Huang N., Bentley J. M. & Hayward M. W. (2012) Soil disturbance by invertebrates in a semi-arid eucalypt woodland: effects of grazing exclusion, faunal reintroductions, landscape and patch characteristics. Proceedings of the Linnean Society of New South Wales. 134, A11-A8.

2011

Hayward, M. W., Hayward, G. J., Tambling, C. J. & Kerley, G. I. H. (2011). Do lions Panthera leo actively select prey or do prey preferences simply reflect responses via evolutionary adaptations to optimal foraging? PLoS ONE 6(8) e23607. 
Hayward, M. W. (2011). Using the IUCN Red List to determine effective conservation strategies. Biodiversity and Conservation 20, 2563-2573.
Hayward, M. W. (2011). Scarcity in the prey community conveys anti-predator benefits. Acta Oecologica 37, 314-320.
Lindsey, P. A., Tambling, C., Brummer, R., Davies-Mostert, H., Hayward, M. W., Marnewick, K., & Parker, D. (2011). Minimum prey and area requirements of cheetahs: implications for reintroductions and management of the species as a managed metapopulation.  Oryx 45, 587-599.
Hayward, M. W., Kowalczyk, R., Krasiński, Z. A., Krasińska, M., Dackiewicz, J. & Cornulier, T. (2011).  Sex allocation in European bison Bison bonasus: has restoration and intensive management altered the evolutionary strategies of an endangered species? Endangered Species Research 15, 53-61.
Burbidge, A. A., Byrne, M., Coates, D., Garnett, S. T., Harris, S., Hayward, M. W., Martin, T. G., McDonald-Madden, E., Mitchell, N. J., Nally, S. & Setterfield, S. (2011). Is Australia ready for assisted colonisation? Policy changes required to facilitate translocation under climate change. Pacific Conservation Biology 17, 259-269. 
Hayward, M. W., Bellchambers, K., Herman, K., Bentley, J. & Legge, S. (2011). Spatial behaviour of yellow-footed rock-wallabies, Petrogale xanthopus, changes in response to active conservation management. Australian Journal of Zoology 59, 1-8.

2010

Hayward, M. W. & Hayward, G. J. (2010). Potential amplification of territorial advertisement markings by black-backed jackals (Canis mesomelas). Behaviour 147, 979-991.
Hayward, M. W. & Zawadzka, B. (2010). Increasing elephant Loxodonta africana density is a more important driver of change in vegetation condition than rainfall. Acta Theriologica 55(4), 289-299.
Bach, L. A., Pedersen, R. P. F., Hayward, M. W., Stagegaard, J. & Loeschske, V., Pertoldi, C. (2010). Assessing re-introductions of the African wild dog (Lycaon pictus) in the Limpopo Valley Conservancy, South Africa, using the stochastic simulation program Vortex.  Journal for Nature Conservation 18 (4), 237-246. 
Hayward, M. W., Hayward, G. J. & Kerley, G. I. H. (2010).  The impact of upgrading roads on the conservation of the threatened flightless dung beetle, Circellum bacchus (F.) (Coleoptera: Scarabaeidae). Coleopterists Bulletin 64 (1), 75-80.

2009

Hayward, M. W. (2009).  The need to rationalize and prioritize threatening processes used to determine threat status in the IUCN Red List.  Conservation Biology 23, 1568-1576.
Hayward, M. W. & Kerley, G. I. H. (2009).  Fencing for conservation: restriction of evolutionary potential or a riposte to threatening processes? Biological Conservation 142, 1-13.
Hayward, M. W. (2009). Conservation management for the past, present and future.  Biodiversity and Conservation 18 (4), 765-775.
Hayward, M. W., Hayward, G. J., Druce, D. & Kerley, G. I. H. (2009) Do fences constrain predator movements on an evolutionary scale? Home range, food intake and movement patterns of large predators reintroduced to Addo Elephant National Park, South Africa. Biodiversity and Conservation 18 (4), 887-899.
Hayward, M. W. & Slotow, R. (2009).  Temporal partitioning of activity in large African carnivores: tests of multiple hypotheses.  South African Journal of Wildlife Research 39 (2), 109-125.
Hayward, M. W. (2009). Bushmeat hunting in the Dwesa and Cwebe Nature Reserves, Eastern Cape, South Africa.  South African Journal of Wildlife Research 39 (1), 70-84.
Hayward, M. W. and Hayward, G. J. (2009). The impact of tourists on lion (Panthera leo) behaviour, stress and energetics. Acta Theriologica 54, 219-224.

2008

Hayward, M. W. & Kerley, G. I. H. (2008). Prey preferences and dietary overlap amongst Africa’s large predators.  South African Journal of Wildlife Research 38(2), 93-108.
Hayward, M.W., de Tores, P. J. and Fox, B.J. (2008).  Post-fire vegetation succession in Agonis linearifolia swamps in the northern jarrah forest of Western Australia.  Conservation Science Western Australia. 7, 35-42
Hayward, M. W. (2008).  Home range overlap in the quokka (Setonix brachyurus) and implications for its mating system.  Conservation Science Western Australia. 7, 57-64
Hayward, M.W. & Munn, A. (2008). Lions, leopards, musk-oxen and marsupials: a light-hearted look at the ups, downs, ins and outs of a postdoctoral career through the eyes of two zoologists.  Australian Zoologist 34, 531-537.

2007

Hayward, M. W., O’Brien, J. & Kerley, G. I. H. (2007).  Carrying capacity of large African predators: predictions and tests. Biological Conservation 139, 219-229.
Hayward, M.W., O’Brien, J. & Kerley, G. I. H. (2007). Testing predictions of the prey of the lion (Panthera leo) derived from modelled prey preferences. Journal of Wildlife Management 71(5): 1567-1575.
Hayward, M.W., Adendorff, J., O’Brien, J., Sholto-Douglas, A., Bissett, C., Moolman, L., Bean, P., Fogarty, A., Howarth, D., Slater, R. & Kerley, G. I. H. (2007).  The reintroduction of large, terrestrial, mammalian predators to the Eastern Cape, South Africa: an assessment. Oryx 41: 205-214.
Hayward, M.W., de Tores, P.J. & Banks, P.B. (2007).  Predicting the occurrence of the quokka, Setonix brachyurus (Macropodidae: Marsupialia), in Western Australia’s northern jarrah forest.  Wildlife Research 34, 194-199.
Hayward, M.W. & Hayward, G.J. (2007). Activity patterns of reintroduced lion Panthera leo and spotted hyaena Crocuta crocuta in Addo Elephant National Park, South Africa.  African Journal of Ecology 45, 135-141.
Hayward, M.W., Adendorff, J., Moolman, L., Dawson, G.J. & Kerley, G.I.H. (2007). The successful reintroduction of leopard Panthera pardus to the Addo Elephant National Park.  African Journal of Ecology 45, 103-104.
de Tores, P.J., Hayward, M.W., Dillon, M.J., & Brazell, R. (2007). Review of the distribution, causes for the decline and recommendations for management of the quokka, Setonix brachyurus (Macropodidae: Marsupialia), an endemic macropodid marsupial from south-west Western Australia. Conservation Science Western Australia 6, 13-73.
Hayward, M.W., Adendorff, J., O’Brien, J., Sholto-Douglas, A., Bissett, C., Moolman, L., Bean, P., Fogarty, A., Howarth, D., Slater, R. & Kerley, G. I. H. (2007).  Practical considerations for the reintroduction of large, terrestrial, mammalian predators based on reintroductions to South Africa’s Eastern Cape Province. The Open Conservation Biology Journal 1, 1-11 doi 10.2174/1874-8392/07

2006

Hayward, M.W., Henschel, P., O’Brien, J., Hofmeyr, M., Balme, G. & Kerley, G.I.H. (2006). Prey preferences of the leopard Panthera pardus. Journal of Zoology 270, 298-313.
Hayward, M.W., O’Brien, J., Hofmeyr, M, & Kerley, G. I. H. (2006). Prey preferences of the African wild dog Lycaon pictus (Canidae: Carnivora): ecological requirements for conservation.  Journal of Mammalogy 87(6), 1122-1131.
Hayward, M.W. (2006). Prey preferences of the spotted hyaena Crocuta crocuta: evidence of dietary competition with the lion Panthera leo.  Journal of Zoology,270, 606-614
Hayward, M.W., O’Brien, J., Hofmeyr, M, & Kerley, G. I. H. (2006). Prey preferences of the cheetah Acinonyx jubatus: morphological limitations or the need to capture rapidly consumable prey before kleptoparasites arrive?  Journal of Zoology,270, 615-627

2005

Hayward, M.W. & Kerley, G.I. (2005). Prey preferences of the lion Panthera leo. Journal of Zoology, 256: 309-322.
Hayward, M.W., de Tores, P.J., Banks, P.B. & Fox, B.J (2005).  Habitat use and preferences of the quokka (Setonix brachyurus Macropodidae: Marsupialia).  Journal of Mammalogy 86(4): 683-688.
Hayward, M. W., White, R. M., Mabandla, K., Bukeye, P. (2005). Larger mammalian fauna of the indigenous forest remnants of the former Republic of Transkei, Eastern Cape, South Africa. South African Journal of Wildlife Research 35(2): 117-124.
Hayward, M. W. (2005).  Diet of the quokka (Setonix brachyurus) (Macropodidae: Marsupialia) in the northern jarrah forest of Australia.  Wildlife Research 32: 15-22.Hayward, M. W., de Tores, P. J., Fox, B. J. and Banks, P. B. (2005).  Using faecal pellet counts along transects to estimate macropodid population density. Wildlife Research 32: 503-507.
Hayward, M.W., de Tores, P.J., Augee, M.J., Fox, B.J & Banks, P.B. (2005).  Survivorship of the quokka (Setonix brachyurus Macropodidae: Marsupialia) in the northern jarrah forest.  Wildlife Research 32, 715-722.

Books

Armstrong, D. P., Hayward, M. W., Moro, D. & Seddon, P. J.  (eds) (2015). Reintroduction Biology in Australia and New Zealand. CSIRO Publishing, Australia. pp 303.

Somers, M. J. & Hayward, M. W. (eds) (2012).  Fencing for Conservation: a restriction of evolutionary potential or a riposte for threatening processes.  Springer, USA.
Hayward, M. W. & Somers, M. J. (eds) (2009).  The Reintroduction of Top-order Predators.  Wiley – Blackwell, London. 

Book chapters

 

Armstrong, D. P., Moro, D., Hayward, M. W., & Seddon, P. J. (2015). The development of reintroduction biology in New Zealand and Australia. In: Reintroduction Biology in Australia and New Zealand pp 1-6 (Eds. Armstrong, D. P., Hayward, M. W., Moro, D. & Seddon, P. J.). CSIRO Publishing, Australia.

Innes, J., Burns, B., Sanders, A. & Hayward, M. W. (2015). The impact of private sanctuary networks on reintroduction programmes. In: ‘Reintroduction Biology in Australia and New Zealand’ pp 185-199 (Eds. Armstrong, D. P., Hayward, M. W., Moro, D. & Seddon, P. J.). CSIRO Publishing, Australia.

Moro, D., Hayward, M. W., Seddon, P. J. & Armstrong, D. P., (2015). Future directions in reintroduction biology of Australian and New Zealand fauna. In: ‘Reintroduction Biology in Australia and New Zealand’ pp 285-290 (Eds. Armstrong, D. P., Hayward, M. W., Moro, D. & Seddon, P. J.). CSIRO Publishing, Australia.

Hayward, M. W., Moseby, K. & Read, J. (2013). The role of predator exclosures in conservation. In: Carnivores of Australia – Past, Present and Future. In press. (Eds. Glen, A. & Dickman, C.). CSIRO, Australia.
Hayward, M. W. (2013). Australia’s south-western woodlands. In: Biomes and Ecosystems: An Encyclopedia. pp. 333-335. (Ed. J. G. Golson). Salem Press, Pasadena, USA.
Hayward, M. W. (2013). Cooper Creek. In: Biomes and Ecosystems: An Encyclopedia. pp 496-498. (Ed. J. G. Golson). Salem Press, Pasadena, USA.
Hayward, M. W. (2013). Kalahari Desert. In: Biomes and Ecosystems: An Encyclopedia. pp 723-726. (Ed. J. G. Golson). Salem Press, Pasadena, USA.
Hayward, M. W. (2013). Namibian savanna woodlands. In: Biomes and Ecosystems: An Encyclopedia. pp 869-871. (Ed. J. G. Golson). Salem Press, Pasadena, USA.
Hayward, M. W. (2013). Simpson Desert. In: Biomes and Ecosystems: An Encyclopedia. pp 1145-1148. (Ed. J. G. Golson). Salem Press, Pasadena, USA.
Allen, B. L., Fleming, P. J. S., Hayward, M. W., Allen, L. R., Engeman, R. M. and Leung, L. K-P. (2012). Top predators as biodiversity regulators: contemporary issues affecting knowledge and management of dingoes in Australia. In: Biodiversity Enrichment in a Diverse World. (Ed. Lameed, G. A.) pp 1-48. Intech, Rijeka, Croatia.
Hayward, M. W. & Somers, M. J. (2012). An introduction to fencing for conservation. In: Fencing for Conservation – Restriction of Evolutionary Potential or a Riposte for Threatening Processes. pp 1-6. (Eds. Hayward, M. W. & Somers, M. J.). Springer-US, New York.
Hayward, M. W. (2012). Perspectives on fencing for conservation based on four case studies: marsupial conservation in Australian forests; bushmeat hunting in South Africa; large predator reintroduction in South Africa; and large mammal conservation in Poland. In: Fencing for Conservation – Restriction of Evolutionary Potential or a Riposte for Threatening Processes. pp 7-20. (Eds. Hayward, M. W. & Somers, M. J.). Springer-US, New York.
Hayward, M. W. (2010). Conservation fencing strategies in Australia. In: The Fence Assessment: A strategic review of fencing policies and impacts in the Great Limpopo Transfrontier Conservation Area and the Kavango-Zambezi Transfrontier Conservation Area. pp 144-148. (Eds. Hanks, J. & Ferguson, K). Wildlife Conservation Society-AHEAD, New York.
Hayward, M. W., Legge, S., Parsons, B., Page, M. & Herman, K. (2010). The reintroduction of woylies (Bettongia penicillata) to Scotia Sanctuary. In: Reintroduction Case Studies 2010.  pp 202-207. IUCN Reintroduction Specialist Group, Gland, Switzerland.
Hayward, M. W. & Somers, M. J. (2009).  The reintroduction of large predators: an introduction and background.  In: The Reintroduction of Top-order Predators. pp 1-9. Blackwell, London.
Hayward, M. W. (2009).  Moving beyond the descriptive: predicting the responses of top-order predators to reintroduction.  In: The Reintroduction of Top-order Predators. pp 345-370 Blackwell, London.
Marnewick, K., Hayward, M. W., Cilliers, D. & Somers, M. J. (2009).  Survival of cheetahs relocated from ranchland to fenced protected areas in South Africa. In: The Reintroduction of Top-order Predators. pp 282-306. Blackwell, London.
Jedrzejewski, W., Hayward, M. W., & Jedrzejewska, B. (2008).  Wolf – moose relationships in the Holarctic zone. In: Proceedings of the VIth International Moose Symposium.  Yakutsk, Russia.