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Dr Craig Shuttleworth

Honorary Visiting Research Fellow

c.shuttleworth@bangor.ac.uk

0000-0003-0076-6789

Dr Craig Shuttleworth

View Dr Craig Shuttleworth’s profile on the Bangor Research Portal

Overview

I have spent thirty years studying the population ecology of red and grey squirrels in Europe and western Canada. My work is currently focussed on pine marten translocation in North Wales.

Additional Contact Information

Mobile 07966150847

Email c.shuttleworth@bangor.ac.uk

Teaching and Supervision

I frequently supervise both undergraduate and postgraduate research, offering local field based studies on Anglesey or in Gwynedd.

Research Interests

My research encompasses viral epidemiological studies with a focus upon Squirrel pox, Adenovirus and Rota virus in sciurid populations; genetic surveillance of red and grey squirrel populations in Europe and North America, wildlife translocation and invasive species management.

I am currently managing the reintroduction of pine marten into North Wales and red squirrel in Clocaenog forest in the North East.

Publications

2025

  • PublishedMitochondrial DNA Analysis of Red Squirrels (Sciurus vulgaris) in North Wales
    O’Meara, D., Shuttleworth, C., Clews-Roberts, R., Everest, D. & Synnott, R., 1 Mar 2025, 14 p. (Evidence Report Series; no. 791)
    Research output: Book/Report › Commissioned report

2024

  • PublishedDeveloping a method for assessing the relative abundance of red squirrels in low-density populations
    Shannon, G., Barton, O., Valle, S. & Shuttleworth, C., 1 Nov 2024, 34 p.
    Research output: Book/Report › Commissioned report
  • PublishedGut bacteria may explain why grey squirrels outcompete reds – new research
    Shuttleworth, C., 20 Feb 2024, The Conversation.
    Research output: Contribution to specialist publication › Article
  • PublishedSignificant differences in the caecal bacterial microbiota of red and grey squirrels in Britain
    Hall, L., Nichols, C., Leng, J., Martelli, F., Shuttleworth, C. & La Ragione, R., 14 Feb 2024, In: Journal of Medical Microbiology. 73, 2
    Research output: Contribution to journal › Article › peer-review

2023

  • PublishedAdenovirus remains widespread in squirrels on Anglesey
    Shuttleworth, C., Everest, D. & McQueen, A., 16 Dec 2023, In: Veterinary Record. 193, 12, p. 490-491 2 p.
    Research output: Contribution to journal › Letter
  • PublishedCapturing red squirrels (Sciurus vulgaris) on camera: A cost-effective approach for monitoring relative abundance and habitat preference
    Shannon, G., Valle, S. & Shuttleworth, C., 3 Oct 2023, In: Ecology and Evolution. 13, 10, e10536.
    Research output: Contribution to journal › Article › peer-review
  • PublishedIndirect effects of pine marten recovery result in benefits to native prey through suppression of an invasive species and a shared pathogen
    Slade, A., White, A., Lurz, P. W. W., Shuttleworth, C., Tosh, D. & Twining, J., 1 Feb 2023, In: Ecological Modelling. 476, 110216.
    Research output: Contribution to journal › Article › peer-review
  • PublishedInvasive genetic rescue: dispersal following repeated culling reinforces the genetic diversity of an invasive mammal
    Synnott, R., Shuttleworth, C., Everest, D., Stevenson-Holt, C., O'Reilly, C., McDevitt, A. & O.Meara, D., Oct 2023, In: Biological Invasions. 25, p. 3323-3339
    Research output: Contribution to journal › Article › peer-review
  • PublishedOpportunistic viral surveillance confirms the ongoing disease threat grey squirrels pose to sympatric red squirrel populations in the UK
    Everest, D., Green, C., Dastjerdi, A., Davies, H., Cripps, R., McKinney, C., Podgornik, G., Stinson, M., O'Hare, S., Bonnie, S., Mill, A., Van der wall, Z., Robinson, N., Trotter, S. & Shuttleworth, C., 20 May 2023, In: Veterinary Record. 192, 10, e2834.
    Research output: Contribution to journal › Article › peer-review
  • PublishedRepeated squirrelpox infection in Welsh red squirrels
    Holmes, J. P., Everest, D. & Shuttleworth, C., 3 Feb 2023, In: Veterinary Record. 192, 3, p. 128-129
    Research output: Contribution to journal › Letter
  • PublishedRestoring our lost wildlife: a review of the ecological, social and economic factors in the potential reintroduction of six mammal species to Cornwall
    Cooper, P., Brazier, R., Puttock, A., Graham, H., Auster, R., Campbell-Palmer, R., Shuttleworth, C. & Pizzi, R., 2023, 258 p.
    Research output: Book/Report › Commissioned report

2022

  • PublishedAn Opportunistic Assessment of the Impact of Squirrelpox Disease Outbreaks upon a Red Squirrel Population Sympatric with Grey Squirrels in Wales
    Shuttleworth, C., Everest, D., Holmes, P., Bell, S. & Cripps, R., 1 Jan 2022, In: Animals. 12, 1
    Research output: Contribution to journal › Article › peer-review
  • PublishedA temporal refuge from predation can change the outcome of prey species competition
    Slade, A., White, A., Lurz, P. W. W., Shuttleworth, C. & Lambin, X., 1 Sept 2022, In: Oikos. Oikos e08565
    Research output: Contribution to journal › Article › peer-review
  • PublishedNon-invasive genetic studies and virus epidemiology
    Everest, D. J., Dastjerdi, A., Davies, H., Shuttleworth, C., Ogden, R., Butler, H. & Sapsford, B., 30 Mar 2022, In: Veterinary Record. 190, p. 204-205
    Research output: Contribution to journal › Article
  • PublishedSquirrelpox outbreak detected in north Wales – without a vaccine, the disease will keep decimating red squirrels
    Shuttleworth, C., 21 Dec 2022, The Conversation.
    Research output: Other contribution

2021

  • PublishedAdenovirus infection in red squirrels
    Shuttleworth, C., Everest, D. J., Stidworthy, M., Halliwell, E. C., Clews-Roberts, R., Paine, V., Bamber, C. & Mason, J., 31 Dec 2021, In: The Veterinary record. 189, p. 444-445
    Research output: Contribution to journal › Article
  • PublishedA practical guide to early surveillance and monitoring
    Robinson, N. J., Everest, D. J., Harry, P. & Shuttleworth, C., 2021, Invasive Alien Species Colonisation Prevention: Your guide to early detection and rapid response . Robinson, N. J. & Shuttleworth, C. M. (eds.). Royal Society of Wildlife Trusts, p. 163-202
    Research output: Chapter in Book/Report/Conference proceeding › Chapter
  • PublishedDisease monitoring and surveillance: case studies in the applied conservation of fragmented red squirrel (Sciurus vulgaris) populations in England and Wales
    Everest, D. J., Floyd, T., Holmes, P., Man, C., Dunnett, C., Locke, R., Savage, L., Sutcliffe, S., Sapsford, B., Shuttleworth, C. & Duff, P., 1 Dec 2021, In: Mammalian Biology. 101, p. 1079-1088
    Research output: Contribution to journal › Article › peer-review
  • PublishedMagical Mammals: Red squirrels in North West Wales
    Shuttleworth, C., 2021, Saving the red squirrel: landscape recovery. Shuttleworth, C. M., Lurz, P. W. W. & Robinson, N. J. (eds.). Red Squirrel Survival Trust, p. 269-278
    Research output: Chapter in Book/Report/Conference proceeding › Chapter
  • PublishedMonitoring escaped captive bred red squirrels in the UK
    Shuttleworth, C., 2021, Saving the red squirrel: landscape recovery. Shuttleworth, C. M., Lurz, P. W. W. & Robinson, N. J. (eds.). Red Squirrel Survival Trust, p. 71-78
    Research output: Chapter in Book/Report/Conference proceeding › Chapter
  • PublishedPreventing grey squirrel establishment on Anglesey
    Shuttleworth, C., 2021, Invasive Alien Species Colonisation Prevention: Your guide to early detection and rapid response . Robinson, N. J. & Shuttleworth, C. M. (eds.). Royal Society of Wildlife Trusts, p. 55-69
    Research output: Chapter in Book/Report/Conference proceeding › Chapter
  • PublishedRecalibrating Risk: Implications of squirrelpox virus for successful red squirrel translocations within mainland UK
    Shuttleworth, C., Brady, D., Cross, P., Gardner, L., Greenwood, A., Jackson, N., McKinney, C., Robinson, N., Trotter, S., Valle, S., Wood, K. & Hayward, M., Feb 2021, In: Conservation Science and Practice. 3, 2, e321.
    Research output: Contribution to journal › Article › peer-review
  • PublishedRed Squirrel Conservation in Clocaenog Forest: Review and Recommendations
    Shuttleworth, C., 2021, 56 p.
    Research output: Book/Report › Commissioned report
  • PublishedSaving the red squirrel: landscape scale recovery
    Shuttleworth, C., Lurz, P. W. W. & Robinson, N., 2021, London: Red Squirrel Survival Trust. 306 p.
    Research output: Book/Report › Book › peer-review
  • PublishedSympatric cases of squirrelpox in red and grey squirrels
    Shuttleworth, C., Everest, D., Wood, C. & Holmes, J. P., 7 May 2021, In: The Veterinary record. 188, 9, p. 358
    Research output: Contribution to journal › Article › peer-review

2020

  • PublishedAre novel ecosystems the only novelty of rewilding?
    Klop-Toker, K., Clulow, S., Shuttleworth, C. & Hayward, M., 1 Nov 2020, In: Restoration Ecology. 28, p. 1318-1320
    Research output: Contribution to journal › Article › peer-review
  • PublishedChallenges and solutions in early detection, rapid response and communication about potential invasive alien species in forests
    de Groot, M., Bullas-Appleton, E., Csóka, G., Csiszár, A., Faccoli, A., Gervasini, A., Kirichenko, N., Shuttleworth, C., Kus Veenvliet, J., O'Hanlon, R., Korda, M., Marinsek, A., Robinson, N., Sweeney, J., Tricarico, E., Verbrugge, L., Williams, D. & Zidar, S., 14 Aug 2020, In: Management of Biological Invasions. 11, p. 637-660
    Research output: Contribution to journal › Article › peer-review
  • PublishedDo differing levels of boldness influence the success of translocation? A pilot study on red squirrels (Sciurus vulgaris)
    Bamber, J. A., Shuttleworth, C. & Hayward, M., 26 Sept 2020, In: Animals. 10, 1748.
    Research output: Contribution to journal › Article › peer-review
  • PublishedEnvisioning the future with 'compassionate conservation': An ominous projection for native wildlife and biodiversity
    Callen, A., Hayward, M. W., Klop-Toker, K., Allen, B. L., Ballard, G., Beranek, C. T., Broekhuis, F., Bugir, C. K., Clarke, R. H., Clulow, J., Clulow, S., Daltry, J. C., Davies-Mostert, H. T., Di Blanco, Y. E., Dixon, V., Fleming, P. J. S., Howell, L. G., Kerley, G. I. H., Legge, S. M. & Lenga, D. J. & 16 others, Major, T., Montgomery, R. A., Moseby, K., Meyer, N., Parker, D. M., Periquet, S., Read, J., Scanlon, R. J., Shuttleworth, C., Tamessar, C. T., Taylor, W. A., Tuft, K., Upton, R. M. O., Valenzuela, M., Witt, R. R. & Wuster, W., Jan 2020, In: Biological Conservation. 241, 108365.
    Research output: Contribution to journal › Article › peer-review
  • PublishedEthical dilemmas when using citizen science for early detection of invasive tree pests and diseases
    Pocock, M. J. O., Marzano, M., Bullas-Appleton, E., Dyke, A., de Groot, M., Shuttleworth, C. & White, R., 1 Nov 2020, In: Management of Biological Invasions. 11, p. 720-732
    Research output: Contribution to journal › Article › peer-review
  • PublishedEvolving grey squirrel management techniques in Europe
    Shuttleworth, C., Robinson, N. J., Halliwell, E. C., Clews-Roberts, R., Peek, H., Podgornik, G., Stinson, M., Rice, S., Finlay, C., McKinney, C., Everest, D. J. & Larsen, K. W., 21 Oct 2020, In: Management of Biological Invasions. 11, 4, p. 747-761
    Research output: Contribution to journal › Article › peer-review
  • PublishedIntroduced Canadian Eastern grey squirrels: squirrelpox virus surveillance and why nothing matters.
    McInnes, C., Shuttleworth, C., Larsen, K. W., Everest, D. J., Bruemmer, C., Carroll, B., Romeo, C., Sainsbury, A. W., Crawshaw, G., Dubois, S., Gillis, L., Gilray, J. & Percival, A., 3 Aug 2020, In: Hystrix-Italian Journal of Mammalogy. 31, p. 95-98
    Research output: Contribution to journal › Article › peer-review
  • PublishedInvasive alien species colonisation prevention: Your guide to early detection and rapid response
    Robinson, N. & Shuttleworth, C., 2020, Newark: Royal Society of Wildlife Trusts. 204 p.
    Research output: Book/Report › Book › peer-review
  • PublishedReinstating trophic cascades as an applied conservation tool to protect forest ecosystems from invasive grey squirrels (Sciurus carolinensis)
    Bamber, J. A., Shuttleworth, C. M., Hayward, M. W. & Everest, D. J., 31 Dec 2020, In: Food Webs. 25, e00164.
    Research output: Contribution to journal › Article › peer-review
  • PublishedResponse to comments on "Compassionate Conservation deserves a morally serious rather than dismissive response - reply to Callen et al., 2020"
    Callen, A., Hayward, M. W., Klop-Toker, K., Allen, B. L., Ballard, G., Beranek, C. T., Broekhuis, F., Bugir, C. K., Clarke, R. H., Clulow, J., Clulow, S., Daltry, J. C., Davies-Mostert, H. T., Di Blanco, Y. E., Dixon, V., Fleming, P. J. S., Howell, L. G., Kerley, G. I. H., Legge, S. M. & Lenga, D. J. & 16 others, Major, T., Montgomery, R. A., Moseby, K., Meyer, N., Parker, D. M., Periquet, S., Read, J., Scanlon, R. J., Shuttleworth, C., Tamessar, C. T., Taylor, W. A., Tuft, K., Upton, R. M. O., Valenzuela, M., Witt, R. R. & Wuster, W., 30 Apr 2020, In: Biological Conservation. 244
    Research output: Contribution to journal › Letter › peer-review

2019

  • PublishedAssessing a potential non-invasive method for viral diagnostic purposes in European squirrels
    Everest, D. J., Tolhurst-Cherriman, D. A. R., Davies, H., Dastjerdi, A., Ashton, A., Blackett, T., Meredith, A. L., Milne, E., Mill, A. & Shuttleworth, C., 4 Jun 2019, In: Hystrix-Italian Journal of Mammalogy. 30, p. 44-50
    Research output: Contribution to journal › Article › peer-review
  • PublishedBritish red squirrels remain the only known wild rodent host for leprosy bacilli
    Schilling, A., Avanzi, C., Ulrich, R. G., Busso, P., Pisanu, B., Ferrari, N., Romeo, C., Mazzamuto, M. V., McLuckie, J., Shuttleworth, C., Del-Pozo, J., Lurz, P. W. W., Escalante-Fuentes, W. G., Ocampo-Candiani, J., Vera-Cabrera, L., Stevenson, K., Chapuis, J.-L., Meredith, A. L. & Cole, S. T., 1 Feb 2019, In: Frontiers in Veterinary Science. 6, p. 8
    Research output: Contribution to journal › Article › peer-review
  • PublishedBy trying to be everything, rewilding risks meaning nothing
    Jachowski, J. D., Bombaci, S., Caravaggi, A., Chalmers, A., Clulow, S., Griffin, A., Hayward, M., Heurich, M., Linnell, J., Marnewick, K., Moehrenschlager, A., Montgomery, A., Shuttleworth, C., Somers, M. & Weise, F., Nov 2019, In: Science. 364, p. 6438
    Research output: Contribution to journal › Letter
  • PublishedControlling grey squirrels in forests and woodlands in the UK
    Gill, R., Ferryman, M., Shuttleworth, C., Lurz, P. W. W., Mill, A., Robertson, P. A. & Dutton, C., 2019, Forest Research. 16 p.
    Research output: Book/Report › Book › peer-review
  • PublishedDeconstructing compassionate conservation
    Hayward, M. W., Callen, A., Allen, B. L., Ballard, G., Broekhuis, F., Bugir, C., Clarke, R. H., Clulow, J., Clulow, S., Daltry, J. C., Davies-Mostert, H. T., Fleming, P. J. S., Griffin, A. S., Howell, L. G., Kerley, G. I. H., Klop-Toker, K., Legge, S., Major, T., Meyer, N. & Montgomery, R. A. & 15 others, Moseby, K., Parker, D. M., Périquet, S., Read, J., Scanlon, R., Seeto, R., Shuttleworth, C., Somers, M. J., Tamessar, C. T., Tuft, K., Upton, R., Valenzuela-Molina, M., Wayne, A., Witt, R. R. & Wuster, W., Aug 2019, In: Conservation Biology. 33, 4, p. 760-768
    Research output: Contribution to journal › Article › peer-review
  • PublishedDetecting viral infection in red squirrels
    Shuttleworth, C., Everest, D. J., Halliwell, E. C., Hulme, B., Wilberforce, L. & Clews-Roberts, R., 30 Apr 2019, In: Veterinary Record. 184, p. 507
    Research output: Contribution to journal › Article
  • PublishedDisease, invasions and conservation: no evidence of squirrelpox virus in grey squirrels introduced to Italy
    Romeo, C., McInnes, C., Dale, T., Shuttleworth, C., Bertolino, S., Wauters, L. A. & Ferrari, N., 1 Feb 2019, In: Animal Conservation. 22, 1, p. 14-23
    Research output: Contribution to journal › Article › peer-review
  • PublishedFirst report of dermatophilosis in wild European red squirrels (Sciurus vulgaris)
    Holmes, P., Everest, D. J., Spiro, S., Wessels, M. & Shuttleworth, C., 30 Sept 2019, In: Veterinary Record Case Reports. 7, 3, p. 3
    Research output: Contribution to journal › Article › peer-review
  • PublishedLeprosy in red squirrels in the UK
    Schilling, A.-K., Del-Pozo, J., Lurz, P. W. W., Stevenson, K., Avanzi, C., Shuttleworth, C., Cole, S. T. & Meredith, A. L., 30 Mar 2019, In: Veterinary Record. 184, p. 416
    Research output: Contribution to journal › Article
  • PublishedPine marten scat holds few clues for squirrel disease
    Shuttleworth, C., Everest, D. J., Dastjerdi, A., Birks, J. & Hayward, M., 7 Dec 2019, In: Veterinary Record. 185, p. 698
    Research output: Contribution to journal › Article
  • PublishedRed squirrel bark-stripping of hornbeam in North Wales
    Shuttleworth, C. & Gill, R., 2019, In: Quarterly Journal of Forestry. 113, 4, p. 275-277
    Research output: Contribution to journal › Article › peer-review
  • PublishedReintroducing rewilding to restoration – Rejecting the search for novelty
    Hayward, M. W., Scanlon, R. J., Callen, A., Howell, L. G., Klop-Toker, K. L., Di Blanco, Y., Balkenhol, N., Bugir, C. K., Campbell, L., Caravaggi, A., Chalmers, A. C., Clulow, J., Clulow, S., Cross, P., Gould, J. A., Griffin, A. S., Heurich, M., Howe, B. K., Jachowski, D. S. & Jhala, Y. V. & 17 others, Krishnamurthy, R., Kowalczyk, R., Lenga, D. J., Linnell, J. D. C., Marnewick, K. A., Moehrenschlager, A., Montgomery, R. A., Osipova, L., Peneaux, C., Rodger, J. C., Sales, L. P., Seeto, R. G. Y., Shuttleworth, C. M., Somers, M. J., Tamessar, C. T., Upton, R. M. O. & Weise, F. J., 1 May 2019, In: Biological Conservation. 233, p. 255-259 5 p.
    Research output: Contribution to journal › Article › peer-review
  • PublishedReleasing grey squirrels into the wild
    Shuttleworth, C., Bertolino, S., Gill, R., Gurnell, J., Hayward, M., Kenward, R. E., Lawton, C., Lurz, P. W. W., McInnes, C. J., Mill, A., Trotter, S. & Wauters, L. A., 31 Mar 2019, In: Veterinary Record. 184, p. 389-390
    Research output: Contribution to journal › Article
  • PublishedRewilding needs to clarify the role of management of invasive species
    Hayward, M., Jachowski, D. S., Shuttleworth, C., Linnell, J. D. C., Allen, B. L., Griffen, A. S., Montgomery, R. A., Caravaggi, A., Weise, F. J., Moehrenschlager, A., Somers, M. J., Clulow, S., Heurich, M. & Marnewick, K., 8 May 2019, In: Science. 364, p. 6438
    Research output: Contribution to journal › Letter
  • PublishedThe search for novelty continues for rewilding
    Hayward, M., Jachowski, J. D., Bugir, C. J., Clulow, J., Krishnamurthy, R., Griffin, A. S., Chalmers, A. C., Linnell, J. D. C., Montgomery, R. A., Somers, M. J., Kowalczyk, R., Heurich, M., Caravaggi, A., Marnewick, K. A., Di Blanco, Y., Shuttleworth, C., Callen, C., Weise, F., Scanlon, R. & Moehrenschlager, A. & 2 others, Howell, L. & Upton, R. M. O., 1 Aug 2019, In: Biological Conservation. 236, p. 584-585
    Research output: Contribution to journal › Article › peer-review
  • PublishedWhy the pine marten is not every red squirrel’s best friend
    Shuttleworth, C. & Hayward, M., 2019, The Conversation.
    Research output: Other contribution

2018

  • PublishedRare woodland wildlife at risk because of 50-year-old tree felling rules
    Shuttleworth, C., 2018, The Conversation.
    Research output: Other contribution
  • PublishedRed squirrel (Sciurus vulgaris) conservation plan for Anglesey & Gwynedd
    Shuttleworth, C. & Halliwell, E. C., 2018, 50 p.
    Research output: Book/Report › Commissioned report
  • PublishedRed squirrels in my garden: guidance and tips to help conserve local populations
    Shuttleworth, C. & Halliwell, E. C., 2018, European Squirrel Initiative. 185 p.
    Research output: Book/Report › Book › peer-review
  • PublishedReducing negative impacts on non-target species during grey squirrel (Sciurus carolinensis) control activities
    Shuttleworth, C., Mill, A. C., Van der Wall, Z., Bertolino, S. & Everest, D. J., 1 Jan 2018, In: Quarterly Journal of Forestry. 112, 1, p. 21-29
    Research output: Contribution to journal › Article › peer-review
  • PublishedThe implications of significant adenovirus infection in UK captive red squirrel (Sciurus vulgaris) collections: How histological screening can aid applied conservation management
    Everest, D. J., Shuttleworth, C., Grierson, S. S., Dastjerdi, A., M.F., S., Duff, J. P., Higgins, R. J., Mill, A. C. & Chantrey, J., 1 Jan 2018, In: Mammalian Biology. 88, p. 123-129
    Research output: Contribution to journal › Article › peer-review
  • PublishedWales should follow Scotland’s lead in protecting the red squirrel’s habitat
    Shuttleworth, C., 2018, Nation Cymru .
    Research output: Other contribution

2017

  • PublishedConfirmation of squirrelpox in Welsh red squirrel
    Everest, D. J., Floyd, T., Donnnacie, B., Irvine, R. M., Holmes, J. P. & Shuttleworth, C., 30 Nov 2017, In: Veterinary Record. 181, p. 514-515
    Research output: Contribution to journal › Article
  • PublishedGrey squirrels are bad for the British countryside – full stop
    Shuttleworth, C., 2017, The Conversation.
    Research output: Other contribution
  • PublishedMathematical models for invasive species management: grey squirrel control on Anglesey
    Jones, H., White, A., Lurz, P. W. W. & Shuttleworth, C., 10 Sept 2017, In: Ecological Modelling. 180, p. 274-284
    Research output: Contribution to journal › Article › peer-review
  • PublishedThe large-scale removal of mammalian invasive alien species in Northern Europe
    Robertson, P. A., Adriaens, T., Lambin, X., Mill, A., Roy, S., Shuttleworth, C. & Sutton-Croft, M., 1 Feb 2017, In: Pest Management Science. 73, p. 273-279
    Research output: Contribution to journal › Article › peer-review
  • PublishedVirus presence within Welsh red squirrels
    Everest, D. J., Holmes, P. & Shuttleworth, C., 30 May 2017, In: The Veterinary record. 180, p. 520-521
    Research output: Contribution to journal › Article

2016

  • PublishedA preliminary comparison of magnum 116 body-grip traps with a live capture trap design during the control of grey squirrels (Sciurus carolinensis)
    Shuttleworth, C., Mill, A. & Van der Waal, Z., Dec 2016, In: International Journal of Pest Management. 63, 4, p. 316-321
    Research output: Contribution to journal › Article › peer-review
  • PublishedEcosystem impacts of an alien invader in Europe, the grey squirrel Sciurus carolinensis
    Gurnell, J., Lurz, P. W. W. & Shuttleworth, C., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 307-329
    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review
  • PublishedExperiences of grey squirrel management in an upland conifer forest
    Halliwell, E. C., Shuttleworth, C., Cartmel, S., Lloyd, I. & Jenkins, R., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 453-474
    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review
  • PublishedGenetic evidence of the effectiveness of grey squirrel control operations: lessons from the Isle of Anglesey
    Signorile, L. & Shuttleworth, C., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 439-452
    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review
  • PublishedGrey squirrel management in Europe - the future
    Shuttleworth, C., Lurz, P. W. W. & Gurnell, J., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative
    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review
  • PublishedGrey squirrel nesting ecology and the use of nest sites in European population management
    Shuttleworth, C., Selonen, V. & Koprowski, J. L., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 349-369
    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review
  • PublishedIdentifying incursion pathways, early detection responses and management actions to prevent grey squirrel range expansion: an island case study in Wales
    Shuttleworth, C., Halliwell, E. C. & Robertson, P. A., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 475-494
    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review
  • PublishedMathematical models of grey squirrel invasion: a case study on Anglesey
    Jones, H. E., White, A., Lurz, P. W. W., Boots, M. & Shuttleworth, C., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 235-255
    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review
  • PublishedRanging behaviour, density and social structure in grey squirrels
    Lawton, C., Shuttleworth, C. & Kenward, R. E., 2016, The grey squirrel: ecology and management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 133-152
    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review
  • PublishedThe grey squirrel: ecology & management of an Invasive Species in Europe
    Shuttleworth, C., Lurz, P. W. W. & Gurnell, J., 2016, European Squirrel Initiative. 505 p.
    Research output: Book/Report › Book › peer-review
  • PublishedThe management of grey squirrel populations in Europe: evolving best practice
    Bertolino, S., Lurz, P. W. W., Shuttleworth, C., Martinoli, A. & Wauters, L. A., 2016, The grey squirrel: ecology and management in Europe . Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 495-516
    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review
  • PublishedThe potential role of ectoparasites in the epidemiology of squirrelpox virus: a possible novel means of intervention to reduce the impact of the disease?
    Cowan, D. P., Mill, A. C., Everest, D. J., McInnes, C. J., Rushton, S. P., Shirley, M. D. F., Start, C. & Shuttleworth, C., 2016, The grey squirrel: ecology and management in Europe . Shuttleworth, C. M., Lurz, P. W. W. & Gurnell, J. (eds.). European Squirrel Initiative, p. 255-289
    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review

2015

  • PublishedAdenovirus detections in juvenile squirrels
    Everest, D. J., Dastjerdi, A., Cowan, D., Gomm, M., Shuttleworth, C., McInnes, C. J., Deane, D., Coulter, L., Mill, A., Rushton, S. P., Jackson, N. L. & Litherland, P., 1 Oct 2015, In: Veterinary Record. 177, p. 373-374
    Research output: Contribution to journal › Article
  • PublishedAssessing causes and significance of red squirrel (Sciurus vulgaris) mortality during regional population restoration: An applied conservation perspective
    Shuttleworth, C., Signorile, , A. L., Everest, D. J., Lurz, P. W. W. & Duff, J. P., 16 Nov 2015, In: Hystrix-Italian Journal of Mammalogy. 26, 2, p. 69-75
    Research output: Contribution to journal › Article › peer-review
  • PublishedBritish red squirrel strongholds: challenges for conservation
    Gurnell, J., Blackett, T., Butler, H., Lurz, P. W. W., Magris, L. & Shuttleworth, C., 2015, Red squirrels: Ecology, Conservation & Management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Hayward, M. W. (eds.). European Squirrel Initiative, p. 211-232
    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review
  • PublishedDeveloping integrated and applied red squirrel conservation programmes: What lessons can Europe learn from a regional grey squirrel eradication programme in North Wales?
    Shuttleworth, C., Schuchert, P., Everest, D. J., McInnes, C. J., Rushton, S. P., Jackson, N. & Kenward, R. E., 2015, Red squirrels: Ecology, Conservation & Management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Hayward, M. W. (eds.). European Squirrel Initiative, p. 233-250
    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review
  • PublishedDeveloping red squirrel re-introduction techniques for use during regional grey squirrel eradication programmes in Europe
    Shuttleworth, C., Jackson, N. & Kenward, R. E., 2015, 2015 Global Reintroduction Perspectives. Soorae, P. (ed.). IUCN/SSC Re-introduction Specialist Group, p. 67-74
    Research output: Chapter in Book/Report/Conference proceeding › Chapter
  • PublishedFuture red squirrel conservation in Clocaenog forest
    Shuttleworth, C. & Hayward, M., 2015, P21018-0026 ed.
    Research output: Book/Report › Commissioned report
  • PublishedIsland Haven: Anglesey red squirrel conservation
    Shuttleworth, C., 2015, Shared Experience of Red Squirrel Conservation Practice . Shuttleworth, C. M., Lurz, P. W. W. & Halliwell, E. C. (eds.). European Squirrel Initiative, p. 151-158
    Research output: Chapter in Book/Report/Conference proceeding › Chapter
  • PublishedPainting the Town Red: Gwynedd red squirrel conservation
    Shuttleworth, C., 2015, Shared Experience of Red Squirrel Conservation Practice . Shuttleworth, C. M., Lurz, P. W. W. & Halliwell, E. C. (eds.). European Squirrel Initiative, p. 159-166
    Research output: Chapter in Book/Report/Conference proceeding › Chapter
  • PublishedRed squirrels: ecology, conservation & management in Europe
    Shuttleworth, C., Hayward, M. & Lurz, P. W. W., 2015, European Squirrel Initiative. 330 p.
    Research output: Book/Report › Book › peer-review
  • PublishedReintroductions and translocations of red squirrels within Europe
    Lawton, C., Waters, C. & Shuttleworth, C., 2015, Red squirrels: Ecology, Conservation & Management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Hayward, M. W. (eds.). European Squirrel Initiative, p. 193-210
    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review
  • PublishedShared experience of red squirrel conservation practice
    Shuttleworth, C., Lurz, P. W. W. & Halliwell, E. C., 2015, European Squirrel Initiative. 220 p.
    Research output: Book/Report › Book
  • PublishedStriving for success: an evaluation of local action to conserve red squirrels (Sciurus vulgaris) in Wales
    Halliwell, E. C., Shuttleworth, C., Wilberforce, L., Denman, H., Lloyd, I. & Cartmel, S., 2015, Red squirrels: Ecology, Conservation & Management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Hayward, M. W. (eds.). European Squirrel Initiative, p. 175-192
    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review
  • PublishedThe impact of adenoviruses on red squirrel (Sciurus vulgaris) conservation – with notes on other enteric viruses of potential significance
    Everest, D. J., Shuttleworth, C., Grierson, S. S., Dastjerdi, A., Stidworthy, M. F., Duff, J. P. & Sainsbury, A. W., 2015, Red squirrels: Ecology, Conservation & Management in Europe. Shuttleworth, C. M., Lurz, P. W. W. & Hayward, M. W. (eds.). European Squirrel Initiative, p. 129-146
    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review

2014

  • PublishedAdenovirus infection: An emerging factor in red squirrel conservation programmes
    Everest, D. J., Shuttleworth, C., Stidworthy, M. F., Grierson, S. S., Duff, P. J. & Kenward, R. E., 1 Oct 2014, In: Mammal Review. 44, 3-4, p. 225-233
    Research output: Contribution to journal › Article › peer-review
  • PublishedAre nest boxes a useful tool in regional red squirrel conservation programs?
    Shuttleworth, C. & Schuchert , P., 15 Dec 2014, In: Hystrix-Italian Journal of Mammalogy. 25, p. 91-94
    Research output: Contribution to journal › Article › peer-review
  • PublishedInter-specific viral infections: Can the management of captive red squirrel collections help inform scientific research?
    Shuttleworth, C., Everest, D. J., McInnes, C. J., Greenwood, A., Jackson, N., Rushton, S. P. & Kenward, R. E., 16 Jun 2014, In: Hystrix-Italian Journal of Mammalogy. 25, 1, p. 18-24
    Research output: Contribution to journal › Article › peer-review
  • PublishedLandscape scale impacts of culling upon a European grey squirrel population: can trapping reduce population size and decrease the threat of squirrel pox virus infection for the native red squirrel?
    Schuchert, P., Shuttleworth, C., McInnes, C. J., Everest, D. J. & Rushton, S. P., 1 Nov 2014, In: Biological Invasions. 16, p. 2381-2391
    Research output: Contribution to journal › Article › peer-review
  • PublishedRed squirrel conservation and disease - Adenovirus in Wales
    Shuttleworth, C., 2014, In: Natur Cymru. 52, p. 41-44
    Research output: Contribution to journal › Article
  • PublishedThe role of spruce in red squirrel (Sciurus vulgaris) conservation in the UK
    Shuttleworth, C. & Lurz, P. W. W., 2014, Forest Ecosystems: Biodiversity, Management and Conservation. Roberts, N. C. (ed.). Nova Science Publishers, p. 63-78
    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review

2013

  • PublishedAdenovirus infection in red squirrels in areas free from grey squirrels
    Everest, D. J., Butler, H., Simpson, V. R., Blackett, T. & Shuttleworth, C., 31 Aug 2013, In: The Veterinary record. 173, 8, p. 199-200
    Research output: Contribution to journal › Article

2012

  • PublishedA systematic assessment of the impact of adenovirus infection on a captive re-introduction project for red squirrels (Sciurus vulgaris)
    Everest, D. J., Shuttleworth, C., Grierson, S. S., Duff, J. P., Jackson, N., Litherland, P. & Kenward, R. E., 31 Aug 2012, In: Veterinary Record. 171, p. 176-184
    Research output: Contribution to journal › Article › peer-review
  • PublishedIntegrating red squirrel (Sciurus vulgaris) habitat requirements with the management of pathogenic tree disease in commercial forests in the UK
    Shuttleworth, C., Lurz, P. W. W., Geddes, N. & Brown, J., 1 Sept 2012, In: Forest Ecology and Management. 279, p. 167-175
    Research output: Contribution to journal › Article › peer-review
  • PublishedRed squirrel population monitoring: Clocaenog forest
    Shuttleworth, C. & Donoyou, T., 2012, IFCW108/001/002 ed.
    Research output: Book/Report › Commissioned report

2010

  • PublishedRetrospective detection by negative contrast electron microscopy of faecal viral particles in wild red squirrels (Sciurus vulgaris) with suspected enteropathy in Great Britain
    Everest, D. J., Stidworthy, M. F., Milne, E. M., Meredith, A. L., Chantery, J., Shuttleworth, C., Blackett, T., Butler, H., Wilkinson, M. & Sainsbury, A. W., 31 Dec 2010, In: Veterinary Record. 167, p. 1007-1010
    Research output: Contribution to journal › Article › peer-review
  • PublishedTurning the grey tide: progress in red squirrel recovery
    Shuttleworth, C., 1 Jun 2010, In: Ecos. 31, p. 27-35
    Research output: Contribution to journal › Article

2009

  • PublishedPCR detection of adenovirus in grey squirrels on Anglesey
    Everest, D. J., Grierson, S. S., Stidworthy, M. & Shuttleworth, C., 31 Oct 2009, In: Veterinary Record. 165, p. 482
    Research output: Contribution to journal › Article › peer-review
  • PublishedThe reintroduction of the red squirrel Sciurus vulgaris to Newborough forest, North Wales: A five year project review
    Shuttleworth, C., Kenward, R. E. & Jackson, N., 2009
    Research output: Book/Report › Commissioned report

2008

  • PublishedAdenovirus-associated deaths in red squirrels on Anglesey
    Everest, D. J., Shuttleworth, C. & Stidworthy, M. F., 31 Oct 2008, In: The Veterinary record. 163, 14, p. 430
    Research output: Contribution to journal › Article › peer-review
  • PublishedRe-introduction of the red squirrel into Newborough forest on the island of Anglesey, UK
    Shuttleworth, C., Kenward, R. E. & Jackson, N., 2008, Global Re-Introduction Perspectives: Re-Introduction Case-Studies from Around the Globe. Soorae, P. S. (ed.). IUCN/SSC Re-introduction Specialist Group, p. 163-166
    Research output: Chapter in Book/Report/Conference proceeding › Chapter

2005

  • PublishedAgainst a grey tide: red squirrel conservation on Anglesey
    Shuttleworth, C., 2005, In: Quarterly Journal of Forestry. 99, p. 213-214
    Research output: Contribution to journal › Article › peer-review
  • PublishedAgainst a grey tide: What future for the red squirrel?
    Shuttleworth, C., 2005, Journal of Practical Ecology & Conservation (Special Series No 4): Living with the Aliens? - Ecology, History, Culture and Management if exotic plants and animals: issues for nature conservation . Rotherham, L. D. (ed.). Vol. 4. p. 87-88
    Research output: Chapter in Book/Report/Conference proceeding › Chapter
  • PublishedGenetic management of the red squirrel, Sciurus vulgaris: a practical approach to regional conservation
    Ogden, R., Shuttleworth, C., McEwing, R. & Cesarini, S., 26 Oct 2005, In: Conservation Genetics. 6, p. 511-525
    Research output: Contribution to journal › Article › peer-review

2004

  • PublishedA study of nest box use by a low density red squirrel (Sciurus vulgaris) population within a commercial conifer plantation
    Shuttleworth, C., 2004, In: Quarterly Journal of Forestry. 98, p. 113-119
    Research output: Contribution to journal › Article › peer-review

2003

  • PublishedA tough nut to crack: red squirrel conservation in Wales
    Shuttleworth, C., 1 Oct 2003, In: Biologist. 50, p. 231-235
    Research output: Contribution to journal › Article › peer-review
  • PublishedRed squirrels on Anglesey
    Shuttleworth, C. & Bailey, M., 2003, In: Natur Cymru. 6, p. 35-37
    Research output: Contribution to journal › Article

2002

  • PublishedRed squirrel conservation on Anglesey, North Wales. A report on the first five years of the Anglesey Red Squirrel Project
    Shuttleworth, C., Bailey, M. & Knott, H., 2002
    Research output: Book/Report › Commissioned report

2001

  • PublishedGrey squirrel control as a tactic in the conservation of the red squirrel: a positive outcome
    Shuttleworth, C., 2001, Advances in Vertebrate Pest Management . Feare, C. J. & Cowan, D. P. (eds.). Filander Verlag, Vol. 3. p. 151-168
    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review
  • PublishedInteractions between the red squirrel (Sciurus vulgaris), great tit (Parus Major) and jackdaw (Corvus monedula) whilst using nest boxes.
    Shuttleworth, C., 1 Oct 2001, In: Journal of Zoology. 255, p. 269-272
    Research output: Contribution to journal › Article › peer-review
  • PublishedThe management of coastal sand dune woodland for red squirrels (Sciurus vulgaris L.)
    Shuttleworth, C. & Gurnell, J., 2001, Coastal Dune Management: Shared Experience of European Conservation Practice. Houston, J., Edmondson, A. S. E. & Rooney, P. J. (eds.). Liverpool University Press, p. 117-127
    Research output: Chapter in Book/Report/Conference proceeding › Chapter
  • PublishedThe use of nest boxes by the red squirrel (Sciurus vulgaris) in conifer plantations
    Shuttleworth, C., 2001, In: Quarterly Journal of Forestry. 95, p. 225-228
    Research output: Contribution to journal › Article › peer-review
  • PublishedTraffic related mortality in a red squirrel (Sciurus vulgaris) population receiving supplemental feeding
    Shuttleworth, C., 1 Jun 2001, In: Urban Ecosystems. 2, p. 109-118
    Research output: Contribution to journal › Article › peer-review

2000

  • PublishedThe foraging behaviour and diet of red squirrels (Sciurus vulgaris) receiving supplemental feeding
    Shuttleworth, C., 1 Sept 2000, In: Wildlife Biology. 6, p. 149-156
    Research output: Contribution to journal › Article › peer-review

1999

  • PublishedThe use of nest boxes by the red squirrel Sciurus vulgaris in a coniferous habitat
    Shuttleworth, C., 1 Mar 1999, In: Mammal Review. 29, p. 61-66
    Research output: Contribution to journal › Article › peer-review

1997

  • PublishedThe effect of supplemental feeding on the diet, population density and reproduction of red squirrels
    Shuttleworth, C., 1997, The Conservation of Red Squirrels (Sciurus vulgaris L.). Gurnell, J. & Lurz, P. W. W. (eds.). Peoples Trust for Endangered Species, p. 13-24
    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review
  • PublishedThe impact of woodland management upon the red squirrel population in Ainsdale National Nature Reserve
    Shuttleworth, C., 1997
    Research output: Book/Report › Commissioned report

1995

  • PublishedRed squirrels (Sciurus vulgaris) on the Sefton Coast: The conservation value of nest boxes
    Shuttleworth, C., 1995, 2nd NPI Red Alert UK Forum for Red Squirrel Conservation. Hughes, D. G. & Tew, T. (eds.). The Federation of Zoological Gardens of Great Britain and Ireland, p. 25-32
    Research output: Chapter in Book/Report/Conference proceeding › Chapter › peer-review

Activities

2022

  • Using remote activated cameras to estimate relative abundance and habitat preference of red squirrels

    Population estimates are essential for the conservation of wildlife. However, accurate estimates can be labour-intensive, particularly for small and medium-sized mammals where capture and handling of individuals is often required. Our study aimed to provide a robust and standardised method of determining red squirrel abundance using remotely activated cameras, while also quantifying how red squirrel abundance relates to key habitat characteristics. Fifty sample points were selected in four habitat categories in forests across Anglesey. We found a strong correlation (r = 0.78) between the number of red squirrels live-trapped over ten-days with the number of camera images recorded during a five-day period. Red squirrel abundance was negatively related to stand canopy openness, while the presence of Scots pine and increased tree species diversity were positively related to squirrel abundance. Our results demonstrate that remotely activated cameras can provide a suitable tool for surveying red squirrel abundance and guiding habitat management.

    19 Dec 2022

    Activity: Oral presentation (Speaker)

2021

  • Opinion piece on forestry law change in Wales

    2021

    Links:

    • https://www.thenational.wales/news/19747937.dr-craig-shuttleworth-wales-forestry-laws-red-squirrels/
    Activity: Types of Public engagement and outreach - Media article or participation (Contributor)

2019

  • Win or bust - Returning the red squirrel to Snowdonia

    One woman's passion for the wilds of North Wales has ignited a resurgence of the charismatic red squirrel. Ecologist Dr Craig Shuttleworth tells us more about the return of red squirrels and their friends the pine martens to Snowdonia National Park.

    3 Jan 2019

    Links:

    • https://www.cnp.org.uk/blog/win-or-bust-returning-red-squirrel-snowdonia
    Activity: Types of Public engagement and outreach - Media article or participation (Contributor)
  • Captive Red Squirrel Management: National Recommendations & Guidance

    Zoological red squirrel collections provide publicly exhibited specimens which are frequently the only opportunity for communities to see this rare native species and to learn about the threats posed to remnant wild populations. Collections are also an important resource for conservation translocation.

    This guidance builds upon historical recommendations for the management and husbandry of captive collections1, evolves the co-ordination of the national studbook and integrates key IUCN guidance2 on the selection of animals for any proposed wild release.

    The maintenance of an effective studbook requires agreement and mutual co-operation. Although this document presents a current best practice, we recognise that management must be adaptive and consequently that the text is a foundation for collaboration and the sharing of future lessons learned and knowledge.

    2019

    Activity: Types of Public engagement and outreach - Media article or participation (Contributor)

Other Grants and Projects

I am currently managing the Heritage Fund funded five year Magical Mammals applied conservation project in north Wales. This aims to widen participation and ldearning within mammal conservation and translocation programmes.

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