I have spent thirty years studying the population ecology of red and grey squirrels in Europe and western Canada. My work is currently focussed on pine marten translocation in North Wales.

Mobile 07966150847

Email c.shuttleworth@bangor.ac.uk

I frequently supervise both undergraduate and postgraduate research, offering local field based studies on Anglesey or in Gwynedd.

My research encompasses viral epidemiological studies with a focus upon Squirrel pox, Adenovirus and Rota virus in sciurid populations; genetic surveillance of red and grey squirrel populations in Europe and North America, wildlife translocation and invasive species management.

I am currently managing the reintroduction of pine marten into North Wales and red squirrel in Clocaenog forest in the North East.

2025

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2019

  • CyhoeddwydAssessing a potential non-invasive method for viral diagnostic purposes in European squirrels
    Everest, D. J., Tolhurst-Cherriman, D. A. R., Davies, H., Dastjerdi, A., Ashton, A., Blackett, T., Meredith, A. L., Milne, E., Mill, A. & Shuttleworth, C., 4 Meh 2019, Yn: Hystrix-Italian Journal of Mammalogy. 30, t. 44-50
    Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
  • CyhoeddwydBritish red squirrels remain the only known wild rodent host for leprosy bacilli
    Schilling, A., Avanzi, C., Ulrich, R. G., Busso, P., Pisanu, B., Ferrari, N., Romeo, C., Mazzamuto, M. V., McLuckie, J., Shuttleworth, C., Del-Pozo, J., Lurz, P. W. W., Escalante-Fuentes, W. G., Ocampo-Candiani, J., Vera-Cabrera, L., Stevenson, K., Chapuis, J.-L., Meredith, A. L. & Cole, S. T., 1 Chwef 2019, Yn: Frontiers in Veterinary Science. 6, t. 8
    Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
  • CyhoeddwydBy trying to be everything, rewilding risks meaning nothing
    Jachowski, J. D., Bombaci, S., Caravaggi, A., Chalmers, A., Clulow, S., Griffin, A., Hayward, M., Heurich, M., Linnell, J., Marnewick, K., Moehrenschlager, A., Montgomery, A., Shuttleworth, C., Somers, M. & Weise, F., Tach 2019, Yn: Science. 364, t. 6438
    Allbwn ymchwil: Cyfraniad at gyfnodolyn › Llythyr
  • CyhoeddwydControlling grey squirrels in forests and woodlands in the UK
    Gill, R., Ferryman, M., Shuttleworth, C., Lurz, P. W. W., Mill, A., Robertson, P. A. & Dutton, C., 2019, Forest Research. 16 t.
    Allbwn ymchwil: Llyfr/Adroddiad › Llyfr › adolygiad gan gymheiriaid
  • CyhoeddwydDeconstructing compassionate conservation
    Hayward, M. W., Callen, A., Allen, B. L., Ballard, G., Broekhuis, F., Bugir, C., Clarke, R. H., Clulow, J., Clulow, S., Daltry, J. C., Davies-Mostert, H. T., Fleming, P. J. S., Griffin, A. S., Howell, L. G., Kerley, G. I. H., Klop-Toker, K., Legge, S., Major, T., Meyer, N., Montgomery, R. A., Moseby, K., Parker, D. M., Périquet, S., Read, J., Scanlon, R., Seeto, R., Shuttleworth, C., Somers, M. J., Tamessar, C. T., Tuft, K., Upton, R., Valenzuela-Molina, M., Wayne, A., Witt, R. R. & Wuster, W., Awst 2019, Yn: Conservation Biology. 33, 4, t. 760-768
    Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
  • CyhoeddwydDetecting viral infection in red squirrels
    Shuttleworth, C., Everest, D. J., Halliwell, E. C., Hulme, B., Wilberforce, L. & Clews-Roberts, R., 30 Ebr 2019, Yn: Veterinary Record. 184, t. 507
    Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl
  • CyhoeddwydDisease, invasions and conservation: no evidence of squirrelpox virus in grey squirrels introduced to Italy
    Romeo, C., McInnes, C., Dale, T., Shuttleworth, C., Bertolino, S., Wauters, L. A. & Ferrari, N., 1 Chwef 2019, Yn: Animal Conservation. 22, 1, t. 14-23
    Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
  • CyhoeddwydFirst report of dermatophilosis in wild European red squirrels (Sciurus vulgaris)
    Holmes, P., Everest, D. J., Spiro, S., Wessels, M. & Shuttleworth, C., 30 Medi 2019, Yn: Veterinary Record Case Reports. 7, 3, t. 3
    Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
  • CyhoeddwydLeprosy in red squirrels in the UK
    Schilling, A.-K., Del-Pozo, J., Lurz, P. W. W., Stevenson, K., Avanzi, C., Shuttleworth, C., Cole, S. T. & Meredith, A. L., 30 Maw 2019, Yn: Veterinary Record. 184, t. 416
    Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl
  • CyhoeddwydPine marten scat holds few clues for squirrel disease
    Shuttleworth, C., Everest, D. J., Dastjerdi, A., Birks, J. & Hayward, M., 7 Rhag 2019, Yn: Veterinary Record. 185, t. 698
    Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl
  • CyhoeddwydRed squirrel bark-stripping of hornbeam in North Wales
    Shuttleworth, C. & Gill, R., 2019, Yn: Quarterly Journal of Forestry. 113, 4, t. 275-277
    Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
  • CyhoeddwydReintroducing rewilding to restoration – Rejecting the search for novelty
    Hayward, M. W., Scanlon, R. J., Callen, A., Howell, L. G., Klop-Toker, K. L., Di Blanco, Y., Balkenhol, N., Bugir, C. K., Campbell, L., Caravaggi, A., Chalmers, A. C., Clulow, J., Clulow, S., Cross, P., Gould, J. A., Griffin, A. S., Heurich, M., Howe, B. K., Jachowski, D. S., Jhala, Y. V., Krishnamurthy, R., Kowalczyk, R., Lenga, D. J., Linnell, J. D. C., Marnewick, K. A., Moehrenschlager, A., Montgomery, R. A., Osipova, L., Peneaux, C., Rodger, J. C., Sales, L. P., Seeto, R. G. Y., Shuttleworth, C. M., Somers, M. J., Tamessar, C. T., Upton, R. M. O. & Weise, F. J., 1 Mai 2019, Yn: Biological Conservation. 233, t. 255-259 5 t.
    Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
  • CyhoeddwydReleasing grey squirrels into the wild
    Shuttleworth, C., Bertolino, S., Gill, R., Gurnell, J., Hayward, M., Kenward, R. E., Lawton, C., Lurz, P. W. W., McInnes, C. J., Mill, A., Trotter, S. & Wauters, L. A., 31 Maw 2019, Yn: Veterinary Record. 184, t. 389-390
    Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl
  • CyhoeddwydRewilding needs to clarify the role of management of invasive species
    Hayward, M., Jachowski, D. S., Shuttleworth, C., Linnell, J. D. C., Allen, B. L., Griffen, A. S., Montgomery, R. A., Caravaggi, A., Weise, F. J., Moehrenschlager, A., Somers, M. J., Clulow, S., Heurich, M. & Marnewick, K., 8 Mai 2019, Yn: Science. 364, t. 6438
    Allbwn ymchwil: Cyfraniad at gyfnodolyn › Llythyr
  • CyhoeddwydThe search for novelty continues for rewilding
    Hayward, M., Jachowski, J. D., Bugir, C. J., Clulow, J., Krishnamurthy, R., Griffin, A. S., Chalmers, A. C., Linnell, J. D. C., Montgomery, R. A., Somers, M. J., Kowalczyk, R., Heurich, M., Caravaggi, A., Marnewick, K. A., Di Blanco, Y., Shuttleworth, C., Callen, C., Weise, F., Scanlon, R., Moehrenschlager, A., Howell, L. & Upton, R. M. O., 1 Awst 2019, Yn: Biological Conservation. 236, t. 584-585
    Allbwn ymchwil: Cyfraniad at gyfnodolyn › Erthygl › adolygiad gan gymheiriaid
  • CyhoeddwydWhy the pine marten is not every red squirrel’s best friend
    Shuttleworth, C. & Hayward, M., 2019, The Conversation.
    Allbwn ymchwil: Cyfraniad arall › Cyfraniad Arall

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2022

  • Using remote activated cameras to estimate relative abundance and habitat preference of red squirrels

    Population estimates are essential for the conservation of wildlife. However, accurate estimates can be labour-intensive, particularly for small and medium-sized mammals where capture and handling of individuals is often required. Our study aimed to provide a robust and standardised method of determining red squirrel abundance using remotely activated cameras, while also quantifying how red squirrel abundance relates to key habitat characteristics. Fifty sample points were selected in four habitat categories in forests across Anglesey. We found a strong correlation (r = 0.78) between the number of red squirrels live-trapped over ten-days with the number of camera images recorded during a five-day period. Red squirrel abundance was negatively related to stand canopy openness, while the presence of Scots pine and increased tree species diversity were positively related to squirrel abundance. Our results demonstrate that remotely activated cameras can provide a suitable tool for surveying red squirrel abundance and guiding habitat management.

    19 Rhag 2022

    Gweithgaredd: Cyflwyniad llafar (Siaradwr)

2021

2019

  • Win or bust - Returning the red squirrel to Snowdonia

    One woman's passion for the wilds of North Wales has ignited a resurgence of the charismatic red squirrel. Ecologist Dr Craig Shuttleworth tells us more about the return of red squirrels and their friends the pine martens to Snowdonia National Park.

    3 Ion 2019

    Cysylltau:

    Gweithgaredd: Mathau o waith ymgysylltu â'r cyhoedd a gwaith maes - Erthygl neu gyfranogiad yn y cyfryngau (Cyfrannwr)
  • Captive Red Squirrel Management: National Recommendations & Guidance

    Zoological red squirrel collections provide publicly exhibited specimens which are frequently the only opportunity for communities to see this rare native species and to learn about the threats posed to remnant wild populations. Collections are also an important resource for conservation translocation.

    This guidance builds upon historical recommendations for the management and husbandry of captive collections1, evolves the co-ordination of the national studbook and integrates key IUCN guidance2 on the selection of animals for any proposed wild release.

    The maintenance of an effective studbook requires agreement and mutual co-operation. Although this document presents a current best practice, we recognise that management must be adaptive and consequently that the text is a foundation for collaboration and the sharing of future lessons learned and knowledge.

    2019

    Gweithgaredd: Mathau o waith ymgysylltu â'r cyhoedd a gwaith maes - Erthygl neu gyfranogiad yn y cyfryngau (Cyfrannwr)

I am currently managing the Heritage Fund funded five year Magical Mammals applied conservation project in north Wales. This aims to widen participation and ldearning within mammal conservation and translocation programmes.