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Dr C Shuttleworth - Staff Profile

Dr C Shuttleworth

Dr C Shuttleworth
Honorary Visiting Research Fellow

Plas Newydd Country House, Llanfairpwll. Anglesey, LL61 6DQ.

Mob: 07966150847 (from U.K.)

Qualifications and Membership of Professional Bodies

BSc (Wildlife Management), PhD (Mammal Ecology)

Scientific Advisor to the Red Squirrel Survival Trust (Patron HRH The Prince of Wales)
Director of Red Squirrels Trust Wales

Research Interests

Pathogenic viral infections in red squirrels
Landscape scale grey squirrel control
Captive breeding and reintroduction of red squirrels
Impact of forest tree disease on red squirrel conservation
Pine Marten reintroduction to northern Gwynedd

Recent Scientific Publications


de Groot M, Bullas-Appleton E, Csóka G, Csiszár A, Faccoli M, Gervasini E, Kirichenko N, ... Shuttleworth CM, ... Kus Veenvliet J (in press) Challenges and solutions in early detection, rapid response and communication about potential invasive alien species in forests.Management of Biological Invasions

Pocock MJO, Marzano M, Bullas-Appleton E, Dyke A, de Groot  M, Shuttleworth CM, Rehema White R (in press) Ethical dilemmas when using citizen science for early detection of invasive tree pests and diseases. Management of Biological Invasions

Shuttleworth CM, Robinson N, Halliwell EC, Clews-Roberts R, Peek H, ... Larsen KW (in press) Evolving grey squirrel management techniques in Europe. Management of Biological Invasions

Callen A, Hayward MW, Klop-Toker K, Allen BL, Ballard G, Beranek CT, ... Shuttleworth CM et al. (2020) Response to: Compassionate Conservation deserves a morally serious rather than dismissive response. Biological Conservation, 244. Doi:10.1016/j.biocon.2020.108517


Gill R, Ferryman M, Shuttleworth C, Lurz P, Mill A Robertson P & Dutton D (2019) Controlling grey squirrels in forests and woodlands in the UK. UK Forestry Standard Technical Note. 16pp. Forest Research

Shuttleworth CM, Everest DJ, Dastjerdi A, Birks JDS and Hayward MW (2019) Pine marten scat holds few clues for squirrel disease. Veterinary Record

Callen A, Hayward M, Tamessar C, Upton R, Broekhuis F, Harriet Davies-Mostert, Ryan Witt, Wolfgang Wuster, Howell L, Katherine Moseby, Guy Ballard, Di Blanco Y, Valenzuela M, Clulow S, Legge S, Major T, Montgomery R, Meyer N, Clulow J, Klop-Toker K, Tuft K, Dixon V, Fleming P, Lenga D, Parker D, Shuttleworth C, Periquet S, Scanlon R, Allen B, Read J, Clarke R, Daltry J, Kerley G, Taylor A (in press) Envisioning the future with ‘Compassionate Conservation’: an ominous projection for biodiversity conservation. Biological Conservation

Shuttleworth CM & Gill R (2019) Red Squirrel Bark-stripping of Hornbeam in North Wales. Quarterly Journal of Forestry 113: 275-277

Hayward MW, Callen A, Allen BL, Ballard G, Broekhuis F, Bugir C, Clarke RH, Clulow J, Clulow S, Daltry JC, Davies-Mostert HT, Fleming PJ, Griffin AS, Howell LG, Kerley GI, Klop-Toker K, Legge S, Major T, Meyer N, Montgomery RA, Moseby K, Parker DM, Périquet S, Read J, Scanlon RJ, Seeto R, Shuttleworth CM, Somers MJ, Tamessar CT, Tuft K, Upton R, Valenzuela‐Molina, M, Wayne A, Witt RR and Wüster W (2019) Deconstructing compassionate conservation. Conservation Biology 33: 760-768. doi:10.1111/cobi.13366

Holmes P, Everest D, Spiro S, Wessels M & Shuttleworth C (in press) First report of dermatophilosis in wild European red squirrels (Sciurus vulgaris)Veterinary Record Case Reports

Hayward MW, Callen A, Allen BL, Ballard G, Broekhuis F, Bugir C, Clarke RH, Clulow J, Clulow S, Daltry, J. C., Davies‐Mostert, H. T., Fleming, P. J., Griffin, A. S., Howell, L. G., Kerley, G. I., Klop‐Toker, K. , Legge, S. , Major, T. , Meyer, N. , Montgomery RA, Moseby K, Parker DM, Périquet S, Read J, Scanlon R, Seeto R, Shuttleworth C, Somers MJ, Tamessar CT, Tuft K, Upton R, Valenzuela‐Molina M, Wayne A, Witt RR & Wüster W (2019), Should the Compassionate Tail Wag the Conservation Dog?. Conservation Biology. doi:10.1111/cobi.13366

Everest DJ, Tolhurst-Cherriman DAR, Davies H, Dastjerdi A, Ashton A, Blackett T, Meredith AL, Milne E, Mill A & Shuttleworth CM (in press) Assessing a potential non-invasive method for viral diagnostic purposes in European squirrels. Hystrix, The Italian Journal of Zoology

Hayward MW, Jachowski D, Bugir CK, Clulow J, Krishnamurthy R, Griffin AS, Chalmers AC, Linnell JDC, Montgomery RA, Somers MJ, Kowalczyk R, Heurich M, Caravaggi A, Marnewick KA, Di Blanco Y, Shuttleworth CM, Callen A, Weise F, Scanlon R, Moehrenschlager A, Howell LG & Upton RMO (2019) The search for novelty continues for rewilding. Biological Conservation (Letter to Editor)

Hayward MW, Jachowski DS, Shuttleworth C, Linnell JDC, Allen BL, Griffin AS, Montgomery RA, Caravaggi A, Weise FJ, Moehrenschlager A, Somers MJ, Clulow S, Heurich M & Marnewick K (2019) Rewilding needs to clarify the role of management of invasive species. eLetter in Science: 364, Issue 6438, eaav5570.

Jachowski D, Bombaci S, Caravaggi A, Chalmers A, Clulow S, Griffin A, Hayward M, Heurich M, Linnell J, Marnewick K, Moehrenschlager A, Montgomery R, Shuttleworth CM, Michael Somers M, Weise F (2019) By trying to be everything, rewilding risks meaning nothing. eLetter in Science: 364, Issue 6438, eaav5570.

Shuttleworth CM, Everest DJ, Halliwell EC, Hulme B, Wilberforce L & Clews-Roberts R (2019) Detecting viral infection in red squirrels. Veterinary Record 184: 507.

Schilling A-K, Del-Pozo J, Lurz PWW,  Stevenson K,  Avanzi C,  Shuttleworth CM,  Cole ST & Meredith AL (2019) Leprosy in red squirrels in the UK. Veterinary Record 184: 416. DOI: 10.1136/vr.l1385

Shuttleworth CM, Bertolino S, Gill R, Gurnell J, Hayward MW, Kenward RE, Lawton C, Lurz PWW, McInnes CJ, Mill A, Trotter S & Wauters LA (2019) Releasing grey squirrels into the wild. Veterinary Record 184, 389-390.

Hayward MW,  Scanlon RJ, Callen Q, Howell LG, Klop-Toker KL, Di Blanco Y, Balkenhol N, Bugir CK, Campbell L, Caravaggi A, Chalmers A, Clulow J, Clulow S, Cross P, Gould JA, Griffin AS, Heurich M, Howe BK, Jachowski DS, Jhala YV, Krishnamurthy R, Kowalczyk R, Lenga DJ, Linnell JDC, Marnewick KA, Moehrenschlager A, Montgomery RA, Osipova L, Peneaux C, Rodger JC, Sales LP, Seeto RGY, Shuttleworth CM, Somers MJ, Tamessar CT, Upton RMO & Weise F (2019) Reintroducing rewilding to restoration – rejecting the search for novelty. Biological Conservation

Schilling AK, Avanzi C, Ulrich RG, Busso P, Pisanu B, Ferrari N, Romeo C, Mazzamuto MV, McLuckie J, Shuttleworth CM et al. (2019) British Red Squirrels Remain the Only Known Wild Rodent Host for Leprosy Bacilli. Veterinary Infectious Diseases.


Romeo C, McInnes C,  Dale T, Shuttleworth CM. Bertolino S, Wauters L, Ferrari N (2018) Disease, invasions and conservation: no evidence of squirrelpox virus in grey squirrels introduced to Italy.  Anim. Conserv. DOI:10.1111/acv.124332017


Everest DJ, Floyd T, Donnnacie B, Irvine RM, Holmes JP & Shuttleworth CM (2017) Confirmation of squirrelpox in Welsh red squirrel. Veterinary Record 181, 514-515.

Everest DJ, Shuttleworth CM, Grierson SS, Dastjerdi A, Stidworthy MF, Duff JP, Higgins RJ, Mill A, Chantrey C (2017) The implications of significant adenovirus infection in UK captive red squirrel (Sciurus vulgaris) collections: How histological screening can aid applied conservation management. Mammalian Biology

Shuttleworth CM, Mill AC, Van Der Waal Z, Bertolino S & Everest DJ (2017) Reducing negative impacts on non-target species during grey squirrel (Sciurus carolinensis) control activities. Quarterly Journal of Forestry

Jones H, White A, Lurz PWW & Shuttleworth CM (2017) Mathematical models for invasive species management: Grey squirrel control on Anglesey. Ecological Modeling 10: 276-284

Everest DJ, Holmes P & Shuttleworth CM (2017) Virus presence within Welsh red squirrels. Veterinary Record 180, 520-521.


Shuttleworth CM, Mill A, Van Der Wall Z (2016) A preliminary comparison of Magnum 116 body-grip traps with a live capture trap design during the control of grey squirrels (Sciurus carolinensis) International Journal of Pest Management

Everest, DJ, Dastjerdi A, Cowan D, Gomm M, Shuttleworth CM, McInnes CJ, Deane D, Coulter L, Mill A, Rushton SP, Jackson NL, Litherland P (2016) Squirrelpox virus antibody detection in juvenile squirrels. Veterinary Record

Robertson, PA, Adriaens T, Lambin X, Mill A, Roy S, Shuttleworth CM, Sutton-Croft M (2016) The large-scale removal of mammalian invasive alien species in northern Europe. Pest Management Science

Shuttleworth CM, Kenward RE, Jackson NJ (2016) Developing red squirrel re-introduction techniques for use during regional grey squirrel eradication programmes in Europe. In: 2015 Global Reintroduction Perspectives. Soorae P (ed.) IUCN/SSC Re-introduction Specialist Group.


Everest, DJ, Dastjerdi A, Cowan D, Gomm M, Shuttleworth CM, McInnes CJ, Deane D, Coulter L, Mill A, Rushton SP, Jackson NL, Litherland P (2015) Adenovirus detections in squirrels from nest boxes. Veterinary Record

Shuttleworth CM, Signorile AL, Everest DJ, Duff JP, Lurz, PWW (2015) Assessing causes and significance of red squirrel (Sciurus vulgaris) mortality during regional population restoration: an applied conservation perspective. Hystrix, The Italian Journal of Zoology.


Shuttleworth, C.M. & Schuchert, P. (2014) Are nest boxes a useful tool in regional red squirrel conservation programs? Hystrix, Italian Journal of Mammalogy doi: 10.4404/hystrix-25.2-10115

Everest, D.J., Shuttleworth, C.M., Stidworthy, M.F., Grierson, S.S., Duff, P.J., Kenward, R.E. (2014) Adenovirus infection: An emerging factor in red squirrel conservation programmes. Mammal Review DOI: 10.1111/mam.12025

Shuttleworth, C.M., Everest, J.D., McInnes, C.J., Greenwood, A., Jackson, N.L., Rushton, S., Kenward, R.E. (2014) Inter-specific viral infections: Can the management of captive red squirrel collections help inform scientific research? Hystrix, Italian Journal of Mammalogy doi:10.4404/hystrix-25.1-10126

Schuchert, P., Shuttleworth, C.M., McInnes, C.J., Everest, D.J., Rushton, S.P. (2014) Landscape scale impacts of culling upon a European grey squirrel population: can trapping reduce population size and decrease the threat of squirrel pox virus infection for the native red squirrel? Biological Invasions doi: 10.1007/s10530-014-0671-8

Edited Books

Robinson N & Shuttleworth CM (2020) Invasive Alien Species Colonisation Prevention: Your guide to early detection and rapid response. 204pp. The Royal Society of Wildlife Trusts. ISBN 978 1 5272 5404 6

Shuttleworth C.M. & Halliwell E.C. (2018) Red squirrels in my garden: Guidance and tips to help encourage and conserve local populations. 120pp. ESI. ISBN 978-0-9547576-5-6

The Grey Squirrel: Ecology & Management of an Invasive Species in Europe. (2016) Eds. Shuttleworth, C.M., Lurz, P.W.W. & Gurnell, J. ESI. ISBN 978-0-954757-62-5 336pp.

Shared Experience of Red Squirrel Conservation Practice. (2015) Eds. Shuttleworth, C.M., Lurz, P.W.W. & Halliwell, E.C. ESI, 220pp.

Red Squirrels: Ecology, Conservation & Management in Europe. (2015) Eds. Shuttleworth C.M., Lurz, P.W.W. & Hayward M.W. ESI, 330pp. ISBN 978-0-9547576-1-8

Recent Online Articles

The Conversation (2019) Why the pine marten is not every red squirrel’s best friend

The Conversation (2018) Rare woodland wildlife at risk because of 50-year-old tree felling rules. 

Nation Cymru (2018) Wales should follow Scotland’s lead in protecting the red squirrel’s habitat.

Campaign for National Parks (2018) Win or bust - Returning the red squirrel to Snowdonia. 

The Conversation (2017) Grey squirrels are bad for the British countryside – full stop.