Professor David Thomas
BSc (Liv), PhD (Liv), CBiol, FSB

Head of School, Chair of Marine Biology and Director of Sêr Cymru National Research Network for Low Carbon, Energy and Environment.


I am Professor in Marine Biology and Head of the School of Ocean Sciences. I was awarded my PhD from the University of Liverpool, UK 1988) for my thesis "Salt tolerances in Cladophora (Chlorophyceae): A study of populations and species". Between 1989 and 1996 I held four research scientist posts in Germany at the University of Bremen, Alfred Wegener Institute, Bremerhaven, University of Oldenburg and Centre for Marine Tropical Ecology, Bremen. I was appointed to my position at Bangor University in July 1996. In 2001 to 2003 I held an Ocean and Climate Research Fellowship at the Hanse Institute for Advanced Study, Germany.

From September 2008 until January 2014 I hold a Research Professor position in the Marine Centre of the Finnish Environment Institute (SYKE) in Helsinki. This position was awarded by the Academy of Finland in their Finland Distinguished Professor Programme (FiDiPro). The theme of the project is "Biogeochemistry of the Baltic Sea in a Changing Climate: From Catchment to Open Sea"

Research Interests

Since 1991 my main research activities have focused on the ecology and biogeochemical processes of sea ice. I have conducted major research expeditions to the Antarctic (1992, 1994, 1997, 1998, 2004/05 and 2006) and Arctic (1993 & 2002). See link. This multidisciplinary work involves collaboration with chemists, glaciologists, physicists and other biologists. These have largely been ship based expeditions, although in 1998 the work was based from a 5 week ice camp in the eastern Weddell Sea, and the 2002 work was a pioneering sea ice study in the White Sea, Russia based from the White Sea Station at Chupa. The Bangor team took part in the ISPOL drift station in the Weddell Sea from November 2004 to March 2005 ( ISPOL and NHM ).
In 2006 we returned to the Weddell Sea as part of the WWOS expedition with Polarstern. Together with colleagues in Finland and Sweden I have also worked on sea ice in the Baltic Sea.

My latest work has focused on the production of, and fate of dissolved organic matter (DOM) within ice. I have participated in large-scale ice tank experiments conducted in ship model testing tanks (EU funded HSVA, Hamburg) to investigate these processes under controlled 'laboratory' conditions, and co-ordinated the experiments in 2001. I also have a more general interest in the role of DOM in aquatic systems, and recent projects involve looking at the role of DOM in northeast Atlantic deep-sea sediments and DOM in freshwaters, estuaries and coastal seas.

My work is not confined to local waters and Polar regions, but I have also conducted fieldwork in the Red Sea, Finland, Vietnam, the Philippines and the North Sea.

I am increasingly interested in the Public Understanding of Science and was awarded a British Association Science and Technology Media Fellowship in 1999. I write regularly for publications with a wider readership including BBC Wildlife Magazine and The Times Higher Education Supplement. I won second prize in the OUP/THES Science Writing Competition 2000, and my work has featured on national and international television and radio. Since 2001 I have been involved in an innovative collaboration with designer-makers Sarah Parker-Eaton and Louise Hibbert in a project called "Plankton Art" (

Research Groups

Research Themes

Research Grants Awarded

2006 to 2009. 'Envirophyte.' European Union Craft Project (with L. Levay). €136,668

2006 to 2007. 'EPS in Antarctic sea ice.' NERC Research Grant (with Underwood, Essex). £64,458

2004 to 2007. 'Antarctic sea ice biogeochemistry.' NERC Research Grant (with H. Kennedy). £256,815

2003 to 2005. 'Biogeochemistry of Antarctic sea ice'. Leverhulme Trust, Research Fellowship. £16,453

2002 to 2004. 'Biogeochemistry of Sea Ice.' Royal Society, Germany - UK Joint Project Grant (with G. Dieckmann,Germany). £10,000

2002 to 2004. 'Antarctic sea Ice.' Royal Society, Research Grant. £7,000

2001 to 2002. 'Using large ice tanks to study the biology and chemistry of sea ice.' NERC Small Research Grant (with H. Kennedy, SOS). £36,012.

2001 to 2003. 'Pathways of organic matter with its implications for biodiversity and sustainable uses in the White Sea (WOMP).' European Union Copernicus Programme. €85,000.

2001 to 2004. 'Dissolved organic matter in coastal ecosystems: transport, dynamics and environmental impact (DOMAINE).' (with P.JleB. Williams & D. Bowers, SOS) European Union, Framework 5 project. €269,836.

1997 to 2000. 'Carbon cycling at the deep ocean boundary layer.' NERC-BENBO thematic programme (with H. Kennedy, SOS). £174,680.

1998 to 1999 'Organic matter production within, and export from Antarctic sea ice platelet layers.' NERC Small Research Grant (with H. Kennedy, SOS) £16,575.

1997 to 1998. 'Biogeochemistry of Antarctic sea ice.' NERC Small Research Grant (with H. Kennedy, SOS) £24,703.

1997-2001. 'Sea Ice Biogeochemistry.' British Council, British German Academic Research Collaboration Programme. (with G.S. Dieckmann, Germany). £8,110.

1997 to 1999. 'Recalcitrant properties of marine dissolved organic matter.' Award to Newly Appointed Science Lecturers. The Nuffield Foundation. £3,500.


Peer Reviewed Journal Papers

Kaartokallio, H, Asmala, E, Autio, R and Thomas, DN. 2016.
Bacterial production, abundance and cell properties in boreal estuaries: Relation to dissolved organic matter quantity and quality. Aquatic Sciences, 78, 525-540.

Räike, A, Kortelainen, P, Mattsson, T and Thomas, DN. 2016.
Long-term trends (1975 - 2014) in the concentrations and export of carbon from Finnish rivers to the Baltic Sea: Organic and inorganic components compared. Aquatic Sciences, 78, 505-523.

Zhou, J, Kotovitch, M, Kaartokallio, H, Moreau, S, Tison, J-L, Kattner, G, Dieckmann, GS, Thomas, DN, Delille, B. 2016.
The impact of dissolved organic carbon and bacterial respiration on pCO2 in experimental sea ice. Progress in Oceanography, doi: 10.1016/j.pocean.2015.12.005

Flemming-Lehtenin V, A Räike, P Kortelainen, P Kaupila and DN Thomas. 2015.
Organic carbon concentration in the northern coastal Baltic Sea between 1975 and 2011. Estuaries and Coasts, 38, 466-481. doi: 10.1007/s12237-0149829-y

Jørgensen, L, CA Stedmon, H Kaartokallio, M Middelboe and DN Thomas. 2015.
Changes in the composition and biovailability of dissolved organic matter during sea ice formation. Limnology and Oceanography, doi: 10.1002/lno.10058

Miller, LA, F Fripiat, BGT Else, JS Bowman, KA Brown, RE Collins, M Ewert, A Fransson, M Gosselin, D Lannuzel, KM Meiners, C Michel, J Nishioka, D Nomura, S Papadimitriou, LM Russell, LL Sørensen, DN Thomas, J-L Tison, MA van Leeuwe, M Vancoppenolle, EW Wolff and J Zhou. 2015.
Methods for biogeochemical studies of Sea Ice: The State of the Art, caveats and recommendations. Elementa - Science of the Anthropocene, doi: 10.12952/journal.elementa.000038

Moreau, S, M Vancoppenole, B DeLille, J-L Tison, J Zhou, M Kotovitch, DN Thomas, N-X Geilfus and H Goosse. 2015.
Drivers of inorganic carbon dynamics in first-year sea ice: a model study. Journal of Geophysical Research - Oceans, doi: 10.1002/2014JC010388

Quintã R, R Santos, DN Thomas and L Le Vay. 2015.
Growth and nitrogen uptake by Salicornia europaea and Aster tripolium in nutrient conditions typical of aquaculture wastewater. Chemosphere,120, 414-421

Quintã, R, P Hill, D. Jones, R. Santos, DN Thomas and L Le Vay. 2015.
Uptake of an amino acid (alanine) and its peptide (trialanine) by the saltmarsh halophytes Salicornia europaea and Aster tripolium and its potential role in ecosystem N cycling and marine aquaculture wastewater treatment. Environmental Engineering, 75, 145-154.

Asmala, E, Bowers, DG, Autio, R, Kaartokallio, H and Thomas, DN. 2014.
Qualitative changes of riverine dissolved organic matter at low salinities due to flocculation. Journal of Geophysical Research-Biogeosciences, 119, 1919-1933.

Asmala, E, R Autio, H Kaartokallio, CA Stedmon and DN Thomas. 2014.
Processing of humic-rich riverine dissolved organic matter by estuarine bacteria: Effects of pre-degredation and inorganic nutrients. Aquatic Sciences, doi: 10.1007/s00027-014-0346-7

Eronen-Rasimus, E, H Kaartokallio, C Lyra, R Autio, H Kuosa, GS Dieckmann and DN Thomas. 2014.
Bacterial community dynamics and activity in relation to dissolved organic matter availability during sea ice formation in a mesocosm experiment. MicrobiologyOpen

Hulatt CJ, H Kaartokallio, E Sonninen, M Oinonen, CA Stedmon and DN Thomas. 2014.
Radiocarbon dating of fluvial organic matter reveals land use impacts in boreal peatlands. Environmental Science and Technology, doi: 10.1021/es5030004

Hulatt, CJ, H Kaartokallio, CA Stedmon, R Autio, E Asmala, E Sonninen, M Oinonen and DN Thomas. 2014.
Bioavailability and radiocarbon age of fluvial dissolved organic matter (DOM) from a northern peatland-dominated catchment: Effect of land-use change. Aquatic Sciences

Papadimitriou S, Kennedy H, Kennedy P, Thomas DN (2014)
Kinetics of ikaite precipitation and dissolution in seawater-derived brines at sub-zero temperatures to 265 K. Geochimica et Cosmochimica Acta 140, 199-211.

Asmala E, DG Bowers, R Autio, H Kaartokallio and DN Thomas. 2014.
Flocculation of riverine humic-rich dissolved organic matter at low salinities. Journal of Geophysical Research-Biogeosciences,doi: 10.1002/2014JG002722

Crabeck O, B Delille, S Rysgaard, DN Thomas, N-X Geilfus, B Else and J-L Tison. 2014.
First 'in situ' determination of gas transport coefficients (DO2, DAr and DN2) from bulk gas concentration (O2, N2, Ar) in natural sea ice. Journal of Geophysical Research-Oceans, doi: 10.1002/2014JC009849

Zhou J, B. Delille, H Kaartokallio, G Kattner, H Kuosa, J-L Tison, R. Autio, GS Dieckmann, K-U Evers, L. Jørgensen, H Kennedy, M Kotovitch, A-M Luhtanen, CA Stedmon and DN Thomas. 2014.
Physical and bacterial controls on the distribution of inorganic nutrients and DOC in sea ice during an experimental ice growth and decay cycle. Marine Chemistry, doi: 10.1016/j.marchem.2014.09.013

DH Søgaard, DN Thomas, S Rysgaard, RN Glud, L Norman, H Kaartokallio, T Juul-Pedersen and NX Geilfus. 2013.
The relative contributions of biological and abiotic processes to the carbon dynamics in subarctic sea ice. Polar Biology, 36, 1761-1777.

H Kaartokallio, DH Søgaard, L Norman, S Rysgaard, B DeLille, JL Tison and DN Thomas. 2013.
Short-term variability in bacteria abundance, cell properties, leucine and thymidine incorporation in subarctic sea ice. Aquatic Microbial Ecology, 71, 57-73.

Papadimitriou S., Kennedy H., Kennedy, D.P Thomas D.N. 2013
Ikaite solubility in seawater-derived brines at 1 atm and sub-zero temperatures to 265 K. Geochimica et Cosmochimica Acta 109 (2013) 241-253

Pieńkowski, A.J., Marret, F., Scourse, J.D. and Thomas, D.N. 2013
Organic-walled microfossils from the north-west Weddell Sea, Antarctica: records from surface sediments after the collapse of the Larsen-A and Prince Gustav Channel ice-shelves. Antarctic Science 25, 565-574.
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Underwood, GJC, Aslam, S, Michel, C, Niemi, A, Norman, L, Meiners, KM, Laybourn-Parry, J, Paterson, H and Thomas, DN. 2013.
Broad-scale predictability of carbohydrates and exopolymers in Antarctic and Arctic sea ice. Proceedings of the National Academy of Sciences of the United States of America, 110, 15734-15739.

E Asmala, R Autio, H Kaartokallio, L Pitkänen, CA Stedmon and DN Thomas. 2013.
Bioavailability of riverine dissolved organic matter in three Baltic Sea estuaries and the effect of catchment land-use. Biogeosciences, 10, 6969-6989.

JM Webb, R Quintã, S Papadimitriou, L Norman, M Rigby, DN Thomas, L Le Vay (2013)
The effect of halophyte planting density on the efficiency of constructed wetlands for the treatment of wastewater from marine aquaculture Ecological Engineering 61, 145-15

M Fischer, DN Thomas, A Krell, G Nehrke, J Goettlicher, L Norman, K Meiners, C Riaux-Gobin and GS Dieckmann. 2013.
Quantification of ikaite in Antarctic sea ice. Antarctic Science, 25, 421-432.

S Müller, A Vähätalo, CA Stedmon, MA Granskog, L Norman, S Aslam, GJC Underwood, GS Dieckmann and DN Thomas. 2013.
Selective incorporation of dissolved organic matter (DOM) during sea ice formation. Marine Chemistry, 155, 148-157.

CJ Hulatt, A-M Lakaneimi, J Puhakka and DN Thomas. 2012.
Energy demands of nitrogen supply in mass cultivation of two commercially important microalgal species, Chlorella vulgaris and Dunaliella tertiolecta. BioEnergy Research, Published online: 2 February 2012. DOI: 10.1007/s12155-011-9175-x
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Meiners, KM, Vancoppenolle, M, Thanassekos, S, Dieckmann, GS, Thomas, DN, Tison, J-L, Arrigo, KR, Garrison, D, McMinn, A, Lannuzel, D, van der Merwe, P, Swadling, K, Smith Jr., WO, Melnikov, I and Raymond, B. 2012.
Chlorophyll a in Antarctic sea ice from historical ice core data. Geophysical Research Letters, 39, L21602, doi:10.1029/2012GL053478

Moate, BD, Bowers, DG and Thomas, DN. 2012.
Measurements of mineral particle absorption properties in turbid estuaries: intercomparison of methods and implications for optical inversions. Estuarine and Coastal Shelf Science, 99 95-107
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Papadimitriou, S, Kennedy, H, Norman, L, Kennedy, P, Dieckmann, GS and Thomas, DN. 2012.
The effect of biological activity, CaCO3 mineral dynamics, and CO2 degassing on the inorganic carbon cycle in sea ice during the late winter-early spring in the Weddell Sea, Antarctica. Journal of Geophysical Research - Oceans, doi: 10.1029/2012JC008058.

SN Aslam, T Cresswell-Maynard, DN Thomas and GJC Underwood. 2012.
Production and characterisation of the intra- and extracellular carbohydrates and polymeric substances (EPS) of three sea ice diatom species, and evidence for a cryoprotective role for EPS. Journal of Phycology, 48, 1494-1509.

A Dausse, A Garbutt, L Norman, LM Jones, S Papadimitriou, PE Robins and DN Thomas. 2012.
Biogeochemical functioning of grazed estuarine tidal marsh along a salinity gradient. Estuarine, Coastal and Shelf Science, 100, 83-92.
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A Räike, P Kortelainen, T Mattsson and DN Thomas. 2012.
36 year trends in dissolved organic carbon export from Finnish rivers to the Baltic Sea. Science of the Total Environment, 435, 188-201.

A-M Lakaniemi, CJ Hulatt, K Wakeham, DN Thomas and JA Puhakka. 2012.
Eukaryotic and prokaryotic microbial communities during microalgal biomass production. Bioresource Technology, 124, 387-393.

CJ Hulatt, A-M Lakaneimi, J Puhakka and DN Thomas. 2012.
Energy demands of nitrogen supply in mass cultivation of two commercially important microalgal species, Chlorella vulgaris and Dunaliella tertiolecta. BioEnergy Research, 5, 669-684.

E Asmala, C Stedmon and DN Thomas. 2012.
Using CDOM absorption coefficients to estimate DOC concentrations in coastal waters. Estuarine, Coastal and Shelf Science, 111, 107-117.

JM Webb, R Quintã, S Papadimitrou, L Norman, M Rigby, DN Thomas and L LeVay. 2012.
Halophyte filter beds for treatment of saline wastewater from aquaculture. Water Research, 46, 5102-5114.

L Tedesco, M Vichi and DN Thomas. 2012.
Process studies on the ecological coupling between sea ice algae and phytoplankton. Ecological Modelling, 226, 120-138.

SN Aslam, GJC Underwood, H Kaartokallio, L Norman, R Autio, M Fischer, GS Dieckmann and DN Thomas. 2012.
Dissolved extracellular polymeric substance (dEPS) production and bacterial growth during sea ice formation in an ice tank study. Polar Biology, 35, 661-676.

Olsen, Y.S., Dausse, A., Garbutt, A., Ford, H., Thomas, D.N. and Jones, D.L. (2011)
Impact of cattle grazing on nitrogen and carbon processing in a temperate saltmarsh. Soil Biology and Biochemistry, 43, 531-541. doi:10.1016/j.soilbio.2010.11.018

CA Stedmon, DN Thomas, S Papadimitriou, MA Granskog and GS Dieckmann. 2011.
Using fluorescence to characterise dissolved organic matter in Antarctic sea ice brines. Journal of Geophysical Research - Biogeosciences, doi:10.1029/2011JG001716

Carver, SM, Hulatt, CJ, Thomas, DN and Tuovinen, OH (2011)
Thermophilic, anaerobic co-digestion of microalgal biomass and cellulose for H2 production. Biodegredation, 22, 805-814. DOI 10.110/s10532-010-9419-z

Hulatt, CJ and Thomas, DN. 2011.
Energy efficiency of an outdoor microalgal photobioreactor sited at mid-temperate latitude. Bioresource Technology, 102, 6687-6695. DOI: 10.1016/j.biortech.2011.03.098

Hulatt, CJ and Thomas, DN. 2011.
Productivity, carbon dioxide uptake and net energy balance of microalgal bubble column photobioreactors. Bioresource Technology, 102, 5775-5787

K.M. Meiners, L.Norman, M.A. Granskog, A. Krell, P. Heil and D. N. Thomas. (2011)
Physico-ecobiogeochemistry of East Antarctic pack ice during the winter-spring transition, Deep- Sea Research II, 58, 1172-1181. doi:10.1016/j.dsr2.2010.10.033

L Norman, CA Stedmon, DN Thomas, M Granskog, S Papadimitriou, KM Meiners, P van der Merwe, D Lanuzel and G.S. Dieckmann.(2011)
Characteristics of dissolved organic matter (DOM) and chromophoric dissolved organic matter (CDOM) in Antarctic sea ice, Deep-Sea Research II, 58, 1075-1091, doi:10.1016/j.dsr2.2010.10.030

S Rysgaard, J Bendtsen, B Delille, GS Dieckmann, R Glud, H Kennedy, J Mortensen, S Papadimitriou, DN Thomas and J-L Tison. 2011.
Sea ice contribution to the air-sea CO2 exchange in the Arctic and Southern Oceans. Tellus B, 63, 823-830.

A-M Lakaniemi, CJ Hulatt, DN Thomas, OH Tuovinen and JA Puhakka. 2011.
Biogenic hydrogen and methane production from Chlorella vulgaris and Dunaliella tertiolecta biomass. Biotechnology for Biofuels, 4, 34 doi:10.1186/1754-6834-4-34

G Massé, ST Belt, X Crosta, S Schmidt, I Snape, DN Thomas and SJ Rowland. 2011.
Highly branched isoprenoind dienes and trienes as proxies for variable sea ice conditions in the Antarctic. Antarctic Science, 23, 487-498.

GJC Underwood, S Fietz, S Papadimitriou, DN Thomas and GS Dieckmann. 2010.
Distribution and properties of dissolved extracellular polymeric substance in Antarctic sea ice habitats. Marine Ecology Progress Series. 404, 1-19. doi: 10.3354/meps08557

GS Dieckmann, G Nehrke, C Uhlig, J Göttlicher, S Gerland, MA Granskog and DN Thomas. 2010.
Ikaite (CaCO_3·6H2O) discovered in Arctic sea ice. The Cryosphere, 4, 227-230

Hulatt, C.J. and Thomas, D.N. (2010)
Dissolved organic matter (DOM) in microalgal photobioreactors: A potential loss in solar energy conversion? Bioresource Technology 101(22) 8690-8697, doi:10.1016/j.biortech.2010.06.086

Sartoris, FJ., Thomas, DN., Cornils, A. and Schnack-Schiel, SB. (2010)
Buoyancy and diapause in Antarctic copepods: The role of ammonium accumulation. Limnology and Oceanography, 55, 1860-1864. doi:10.4319/lo.2010.55.5.1860

Hulatt, C, DN Thomas, DG Bowers, L Norman and C Zheng. 2009.
Exudation and decomposition of coloured dissolved organic matter (CDOM) from some macroalgae. Estuarine and Coastal Shelf Science. 84, 147-153

Meiners, K.M., Papadimitriou, S., Thomas, D.N., Norman L. and Dieckmann G.S. 2009.
Biogeochemical conditions and ice algal photosynthetic parameters in Weddell Sea ice during early spring. Polar Biology, doi:10.1007/s00300-009-0605-6

Papadimitriou, S., Thomas, D.N., Kennedy, H., Kuosa, H., and Dieckmann, G.S. 2009.
Inorganic carbon removal and isotopic enrichment in Antarctic sea ice gap layers during early austral summer. Marine Ecology Progress Series. doi:10.3354/meps08049

Pieńkowski, A.J., Marret, F., Thomas, D.N., Scourse, J.D. and Dieckmann, G.S. 2009.
Dinoflagellates in a fast -ice covered inlet of the Riiser-Larsen Ice Shelf (Antarctica): Polar Biology 32, 1331-1343. doi:10.1007/s00300-009-0630-5

Van der Merwe, P., Lannuzel, D., Mancuso Nichols, C.A., Meiners, K., Heil, P., Norman, L., Thomas, D.N. and Bowie, A.R. 2009.
Biogeochemical observations during the winter-spring transition in East Antarctic sea ice: Evidence of iron and exopolysaccharide controls. Marine Chemistry. 115, 163-175
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Schnack-Schiel, SB, C Haas, J Michels, E Mizdalski, H Schünemann, M Steffens, DN Thomas. 2008.
Copepods in sea ice of the western Weddell Sea during austral spring 2004. Deep Sea Research II. 55, 1056- 1067.

Dieckmann, GS, Nehrke, G, Papadimitriou, S, Göttlicher, J., Steininger, R., Kennedy, H., Wolf-Gladrow, D. and Thomas, DN. 2008.
Calcium carbonate as ikaite crystals in Antarctic ice. Geophysical Research Letters Vol. 35. doi:10.1029/2008GL033540

Tison, J-L, T Worby, B deLille, F Brabant, S Papadimitriou, DN Thomas, J de Jong, D Lannuzel and C Haas. 2008
Temporal evolution of decaying summer first-year sea ice in the Western Weddell Sea. Deep Sea Research II. 55, 975-987

Clarke, A.C., MP Meredith, MI Wallace, M Brandon and DN Thomas. 2008
Seasonal and interannual variability in temperature, chlorophyll and macronutrients in Ryder Bay, northern Marguerite Bay, Antarctica. Deep Sea Research II. 55, 1988-2006.

Michels, J., GS Dieckmann, DN Thomas, S Schnack-Schiel, A Krell, P Assmy, H Kennedy, S Papadimitriou and B Cisewski. 2008
Short term biogenic particle flux under late spring sea ice in the western Weddell Sea. Deep Sea Research II. 55, 1024-1039.

Zemmelink, H.J., L Houghton, JWH Dacey, J Stefels, BP Koch, G Schröder, M Wisotzki, A Scheltz, DN Thomas, S Papadimitriou, H Kennedy, H Kuosa and T Dittmar. 2008
Stratification and the distribution of phytoplankton, nutrients, inorganic carbon, and sulfur in the surface waters of the Weddell Sea. Deep Sea Research II. 55, 988-999.

Stedmon C., Thomas D.N., Granskog M., Kaartokallio H., Papadimitriou S. and Kuosa H. 2007
The characteristics of dissolved organic matter in Baltic coastal sea ice: Allochthonous or autochthonous origin? Environmental Science and Technology. DOI: 10.1021/es071210f

Evans, C.D., C Freeman, LG Cork, DN Thomas, B Reynolds, MF Billet, MH Garnett and D Norris. 2007
Evidence against recent climate-induced destabilisation of soil carbon from 14C analysis of riverine dissolved organic matter. Geophysical Research Letters. 34, L07407, doi 10.1029/2007GL029431
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Kaartokallio, H., H Kuosa, DN Thomas, MA Granskog and K Kivi. 2007
Biomass, composition and activity of organism assemblages along a salinity gradient in sea ice subjected to river discharge in the Baltic Sea. Polar Biology.30, 183-197.
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Papadimitriou, S., DN Thomas, H Kennedy, C Haas, H Kuosa, A Krell and GS Dieckmann. 2007
Biogeochemical composition of natural sea ice brines from the Weddell Sea during early austral summer. Limnology and Oceanography. 52 1809-1823
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Granskog, M.A., Kaartokallio, H., Kuosa, H., Thomas, D.N. and Vainio, J. 2006
Sea ice in the Baltic Sea - A Review. Estuarine and Coastal Shelf Science, 70, 145-160.
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Steffens, M., MA Granskog, H Kaartokallio, H Kuosa, K Luodekari, S Papadimitriou and DN Thomas. 2006
Spatial variation of biogeochemical properties of landfast sea ice in the Gulf of Bothnia (Baltic Sea). Annals of Glaciology. 44, 80-87.
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Granskog, M.A., H Kaartokallio, DN Thomas and H Kuosa. 2005
Influence of freshwater inflow on the inorganic nutrient and dissolved organic matter within coastal sea ice and underlying waters in the Gulf of Finland (Baltic Sea). Estuarine and Coastal Shelf Science. 65, 109-122.
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Granskog, M.A., Kaartokallio, H., Kuosa, H., Thomas, D.N., Ehn, J. and Sonninen, E. 2005
Scales of horizontal patchiness in chlorophyll-a, chemical and physical properties of landfast sea ice in the Gulf of Finland (Baltic Sea). Polar Biology, 28, 276- 283.
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Krell, A., Schnack-Schiel, S.B., Thomas, D.N., Kattner, G., Zipan, W. and Dieckmann, G.S. 2005
Phytoplankton dynamics in relation to hydrography, nutrients and zooplankton at the onset of sea ice formation in the eastern Weddell Sea (Antarctica). Polar Biology, 28, 700-713.
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Mock, T. and Thomas, D.N.. 2005
Sea ice - Recent advances in microbial studies Environmental Microbiology. 7, 605-619.
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Thomas, D.N. 2005
Photosynthetic microbes in freezing deserts. Trends in Microbiology, 13, 87-88.
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Papadimitriou, S, Kennedy, H, Kattner, G, Dieckmann, GS and Thomas, DN 2004
Experimental evidence for carbonate precipitation and CO2 degassing during sea ice formation. Geochimica et Cosmochimica Acta., 68, 1749-1761.
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Arrigo, K.R. and Thomas, D.N. 2004
The importance of sea ice for the Southern Ocean ecosystem. Antarctic Science, 16, 471-486.
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Bowers, D.G., Evans, D., Thomas, D.N., Ellis, K. and Williams, P.J.leB. 2004
Interpreting the colour of an estuary. Estuarine Coastal and Shelf Science, 59, 13-20.
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Granskog, M.A., Virkkunen, K., Thomas, D.N., Ehn, J., Kola, H. and Martma, T. 2004
Chemical properties of brackish water ice in the Bothnian Bay, the Baltic Sea. Journal of Glaciology, 50, 292-302.

Kattner, G., Thomas, D.N., Haas, C., Kennedy, H. and Dieckmann, G.S. 2004
Surface ice and gap layers in Antarctic sea ice: highly productive habitats. Marine Ecology Progress Series, 277, 1-12,
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Papadimitriou, S., Kennedy, H. and Thomas, D.N. 2004
Rates of organic carbon oxidation in deep sea sediments in the eastern North Atlantic from pore water profiles of O2 and δ13C of dissolved inorganic carbon. Marine Geology, 212, 97-111.
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Patsayeva, S., R Reuter and DN Thomas. 2004
Fluorescence of dissolved organic matter in seawater at low temperatures and during ice formation. EARSeL eProceedings 3, 227-238.

Schnack-Schiel, S.B., Kattner, G., Dieckmann, G.S. and Thomas, D.N. 2004
Copepods under summer fast ice in the eastern Weddell Sea, Antarctica. Polar Biology, 27, 502- 506.

Thomas, D.N.. 2003
Iron limitation in the Southern Ocean. Science 302, 565.

Krell, A., C Ummenhofer, G. Kattner, A Naumov, DW Evans, GS Dieckmann and DN Thomas. 2003
The biology and chemistry of land fast-ice in the White Sea, Russia - A comparison of winter and spring conditions. Polar Biology. 26, 707-719.
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Mock, T., GS Dieckmann, C Haas, J-L Tison, A Krell, AL. Belem, S. Papadimitriou and DN Thomas. 2002
Micro-optodes in a sea ice: A new approach to investigate oxygen dynamics during sea ice formation. Aquatic Microbial Ecology 29, 297-306.
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Thomas, D.N. and Dieckmann, G.S. 2002
Biogeochemistry of Antarctic sea ice. Oceanography and Marine Biology: an Annual Review. 40, 143 -169.

Brierley, A.S. and DN Thomas. 2002
The ecology of Southern Ocean pack ice. Advances in Marine Biology 43, 171-278.

Kennedy, H., Thomas, D.N., Kattner, G., Haas, C. and Dieckmann, G.S. 2002
Particulate organic carbon in Antarctic summer sea ice: Concentration and stable carbon isotopic composition. Marine Ecology Progress Series 238, 1-13.
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Papadimitriou, S., Kennedy, H., Bentaleb, I. and Thomas, D.N.. 2002
Dissolved organic carbon in sediments from the NE Atlantic Continental Slope. Marine Chemistry 79, 37-47.
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Post, A.F., Z Dedej, R. Gottlieb, H Li, DN Thomas, M El-Absawi, A El-Naggar, M El-Gharabawi and U Sommer. 2002
Spatial and temporal distribution of Trichodesmium spp. in the stratified Gulf of Aqaba, Red Sea. Marine Ecology Progress Series 239, 241-250.
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Thomas, D.N. and Dieckmann, G.S. 2002
Antarctic sea ice - a habitat for extremophiles. Science 295. 641-644.
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Thomas, D.N., H Kennedy, G Kattner, D Gerdes, C Gough and GS Dieckmann. 2001
Biogeochemistry of platelet ice: Influence on particle flux under land fast sea ice during summer at Drescher Inlet, Weddell Sea, Antarctica. Polar Biology. 24. 486-496. (Also published in 2002. In: Ecological Studies in the Antarctic sea ice Zone (Eds. WE Arntz and A Clarke). pp169-179. Springer Verlag Berlin Heilelberg).
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Thomas, D.N., R. Engbrodt, V. Giannelli, G. Kattner, H. Kennedy, C. Haas and GS Dieckmann. 2001
Dissolved organic matter in Antarctic sea ice. Annals of Glaciology 33. 297-303.

Giannelli, V., Thomas, D.N., Haas, C., Kattner, G., Kennedy, H.A. and Dieckmann, G.S. 2001
Behaviour of dissolved organic matter and inorganic nutrients during experimental sea ice formation. Annals of Glaciology, 33, 317–321.

Haas, C., Thomas, D.N. and Bareiss, J. 2001
Surface properties and processes of perennial Antarctic sea ice in summer. Journal of Glaciology, 47, 613-625.

Herborg, L.M., Thomas, D.N., Kennedy, H., Haas, C. and Dieckmann, G.S. 2001
Dissolved carbohydrates in Antarctic sea ice. Antarctic Science, 13, 119-125.
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Schnack-Schiel, S.B., Dieckman, G.S., Gradinger, R., Melnikov, I.A., Spindler, M. and Thomas, D.N. 2001
Meiofauna in sea ice of the Weddell Sea (Antarctica). Polar Biology, 24, 724-728.

Schnack-Schiel, S.B., Thomas, D.N., Haas, C., Dieckmann, G.S. and Alheit, R. 2001
The occurrence of the copepods Stephos longipes (Calanoida) and Drescheriella glacialis (Harpacticoida) in summer sea ice in the Weddell Sea. Antarctic Science, 13, 150-157.
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Haas, C., F Cottier, LH Smedsrud, DN Thomas, U Buschmann, D Dethleff, S Gerland, V Giannelli, J Hoelemann, J-L Tison and P Wadhams. 1999
Multidisciplinary ice tank study shedding new light on sea ice growth processes. EOS, Transactions, American Geophysical Union 80. 507-513.

Thomas, D.N., RJ Lara, C Haas, SB Schnack-Schiel, E-M Nöthig, GS Dieckmann, G. Kattner and E. Mizdalski. 1998
Biological soup within decaying summer sea ice in the Amundsen Sea, Antarctica. In Antarctic Sea Ice Biological Processes, Interactions and Variability (eds. M Lizotte and K Arrigo). American Geophysical Union. Antarctic Research Series 73. 161 - 171.

Dieckmann, G.S., H Eicken, C Haas, DL Garrison, M Gleitz, M Lange, E-M Nöthig, M Spindler, CW Sullivan, DN Thomas and J Weissenberger. 1998
A compilation of data on sea ice algal standing crop from the Bellingshausen, Amundsen and Weddell Seas from 1983 to 1994. In Antarctic Sea Ice Biological Processes, Interactions and Variability (eds. M Lizotte and K Arrigo). American Geophysical Union. Antarctic Research Series 73. 85 - 92.

Lara, R.J, V Rachold, G Kattner, H-W Hubberten, G Guggenberger, A Skoog and DN Thomas. 1998
Dissolved organic matter and nutrients in the Lena River, Siberian Arctic: characteristics and distribution. Marine Chemistry 59. 301-310.

Skoog, A., DN Thomas, RJ Lara and K-U Richter. 1997
Methodological investigations on DOC determinations by the HTCO method. Marine Chemistry 56. 39-44.

Lara, R.J., U Hubberten, DN Thomas, MEM Baumann and G Kattner. 1997
Dissolved organic matter studies in enclosed systems: Application of hydrophobic fractionation for the assessment of organic nitrogen dynamics. Journal of Marine Systems 13. 155-161.

Weykam, G., Thomas, D.N. and Wiencke, . 1997
Growth and photosynthesis of the Antarctic red algae, Palmaria decipiens (Palmariales) and Iridaea cordata (Gigartinales) during and following extended periods of darkness. Phycologia 36. 395-405.

Gómez, I., Wiencke, C. and Thomas, D.N. 1996
Age-dependent variations in photosynthetic characteristics of the Antarctic brown alga Ascoseira mirabilis Skottsberg. European Journal of Phycology 31. 167-172.

Peters, E. and Thomas, D.N. 1996
Prolonged darkness and diatom mortality 1: Marine Antarctic species. Journal of Experimental Marine Biology and Ecology. 207. 25-41.

Peters, E. and Thomas, D.N. 1996
Prolonged nitrate exhaustion and diatom mortality - A comparison of polar and temperate Thalassiosira species. Journal of Plankton Research 18. 953-968.

Thomas, D.N. and Lara, R.J. 1995
Photodegradation of algal dissolved organic carbon. Marine Ecology Progress Series 116. 309-310.

Thomas, D.N., RJ Lara, H Eicken, G Kattner and A Skoog. 1995
Dissolved organic matter in Arctic multi- year sea ice during winter: Major components and relationship to ice characteristics. Polar Biology 15. 477-483.

Gómez, I., Thomas, D.N. and Wiencke, C. 1995
Longitudinal profiles of growth, photosynthesis and light independent carbon fixation in the Antarctic brown alga Ascoseira mirabilis. Botanica Marina 38. 157-164.

Gleitz, M., M Rutgers vd Loeff, DN Thomas, GS Dieckmann and FJ Millero. 1995
Comparison of summer and winter inorganic carbon, oxygen and nutrient concentrations in Antarctic sea ice brine. Marine Chemistry 51. 81-91.
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Haas, C. and Thomas, D.N. 1995
Glacial-ice fragments in Antarctic sea ice. Journal of Glaciology 41. 432-435.

Lara, R.J. and Thomas, D.N. 1995
Formation of recalcitrant organic matter: Humification dynamics of algal derived dissolved organic carbon and its hydrophobic fractions. Marine Chemistry 51. 193-199.

Schnack-Schiel, S.B., DN Thomas, GS Dieckmann, H Eicken, R Gradinger, M Spindler, J Weissenburger, E Mizdalski and K Beyer. 1995
Life cycle strategy of the Antarctic calanoid copepod Stephos longipes. Progress in Oceanography 36. 45-75.

Lara, R.J. and Thomas, D.N. 1994
XAD-fractionation of "new" dissolved organic matter: Is the hydrophobic fraction seriously underestimated? Marine Chemistry 47. 93-96.

Lara, R.J. and Thomas, D.N. 1994.
Isolation of Marine Dissolved Organic Matter: Evaluation of Sequential Combinations of XAD Resins 2,4 and 7. Analytical Chemistry 66. 2417-2419.

Thomas, D.N. and Gleitz, M. 1993
Allocation of photoassimilated carbon into major algal metabolite fractions: Variation between two diatom species isolated from the Weddell Sea (Antarctica). Polar Biology 13. 281-286.

Gleitz, M. and Thomas, D.N. 1993
Variation in phytoplankton standing stock, chemical composition and physiology during sea ice formation in the southeastern Weddell Sea, Antarctica. Journal of Experimental Marine Biology and Ecology 173. 211-230.

Thomas, D.N., MEM Baumann and M Gleitz. 1992
Efficiency of carbon assimilation and photoacclimation in a small unicellular Chaetoceros species from the Weddell Sea (Antarctica): Influence of temperature and irradiance. Journal of Experimental Marine Biology and Ecology 157. 195-209.

Gleitz, M. and Thomas, D.N. 1992
Physiological responses of a small Antarctic diatom (Chaetoceros sp.
to simulated environmental contraints associated with sea-ice formation. Marine Ecology Progress Series 88. 271-278.

Thomas, D.N. and Kirst, G.O. 1991
Differences in osmoacclimation between sporophytes and gametophytes of the brown alga Ectocarpus siliculosus. Physiologia Plantarum 83. 281-289.

Thomas, D.N. and Kirst, G.O. 1991
Salt tolerance of Ectocarpus siliculosus (Dillw.) Lyngb: Comparison of gametophytes, sporophytes and isolates of different geographic origin. Botanica Acta 104. 26-36.

Thomas, D.N. and Wiencke, C. 1991
Photosynthesis, dark respiration and light independent carbon fixation of endemic Antarctic macroalgae. Polar Biology 11. 329-337.

Karsten, U., DN Thomas, G Weykam, C Daniel and GO Kirst. 1991
A simple and rapid method for extraction and separation of low molecular weight carbohydrates from macroalgae using high-performance liquid chromatography. Plant Physiology and Biochemistry 29. 373-378.

Thomas, D.N., JC Collins and G Russell. 1990
Inter- population differences in the salt tolerances of two Cladophora species. Estuarine, Coastal and Shelf Science 30. 201-206.

Thomas, D.N., Collins, J.C. and Russell, G. 1989
Physiological responses to salt stress of two ecologically different Cladophora species. Botanica Marina 32. 259- 265.

Thomas, D.N., Collins, J.C. and Russell, G. 1988
Interactive effects of temperature and salinity upon net photosynthesis of Cladophora glomerata (L.) Kutz. and C. rupestris (L.) Kutz. Botanica Marina 31. 73-77.


Williams PJ LeB, Evans D, Roberts DJ, Thomas DN. 2015
Art Forms from the Abyss Ernst Haeckel's Images from the HMS Challenger Expedition. Prestel ISBN: 978-3-7913-8141-1
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DN Thomas and DG Bowers. 2012. Introducing Oceanography. 151 pages. Dunedin Academic Press Ltd. ISBN 9781780460017.
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MJ Kaiser, M Attrill, S Jennings, DN Thomas, D Barnes, AS Brierley, JG Hiddink, H Kaartokallio, N Polunin and D Raffaelli. 2011.
Marine Ecology - Processes, Systems and Impacts. 2nd Edition. 576 pages. Oxford University Press. ISBN 9780199227020.

Thomas, DN and Dieckmann, GS(Eds.). 2010.
Sea Ice - 2nd Edition. Wiley-Blackwell Publishing, Oxford. 600 pages. ISBN 978-1-4051-8580-6
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Thomas, D.N., GE Fogg, P Convey, C Fritsen, J-M Gilli, R Gradinger, J Laybourn-Parry, K Reid, D Walton. 2008
Biology of Polar Regions. 416 pages. Oxford University Press. ISBN-13:9780199298136
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Thomas, D.N. 2007
Surviving Antarctica. Natural History Museum. 96 pages. ISBN 0565092170. Also published by Firefly Books (USA and Canada) ISBN 1554072941 and Museum Victoria (Australia and New Zealand).
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Kaiser, M.J., M Attrill, S Jennings, DN Thomas, D Barnes, AS Brierley, N Polunin, D Raffaelli and P J LeB Williams. 2005
Marine Ecology - Processes, Systems and Impacts. Oxford University Press. 584 pages. ISBN 019924975X.
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Søndergaard, M. and Thomas, D.N. (Eds.) 2004
Dissolved Organic Matter (DOM) in Aquatic Ecosystems: A Study of European Catchments and Coastal Waters. 74 pages. ISBN 8789143256

Thomas, D.N. 2004
Frozen Oceans - The Floating World of Pack Ice. Natural History Museum, London. 240 pages. ISBN 0565 091883. Also published by Firefly Books (USA and Canada) ISBN 1554070007 and CSIRO (Australia and New Zealand) ISBN 0643090878
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Thomas, D.N.2002
Seaweeds. Natural History Museum. 96 pages. ISBN 0565091751. Also published by the Smithsonion Institute Press (USA and Canada) ISBN 1-58834-050-3 and CSIRO (Australia and New Zealand) ISBN 0565091751
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Williams P.J. LeB, Thomas, D.N. and Reynolds, C.S. (Eds.) 2002
Phytoplankton Productivity - Carbon assimilation in marine and freshwater ecosystems. Blackwell Publishing. 320 pages. ISBN 0632057114.

Thomas, D.N. and Dieckmann, G.S. (Eds.) 2003
Sea ice - An Introduction to its Physics, Chemistry, Biology and Geology. Blackwells Publishing, Oxford. 416 pages. ISBN 0632058080. Contains - DN Thomas and S. Papadimitriou. Chapter 9, Biogeochemistry of sea ice. Pages 267-302.
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Book Reviews

DN Thomas. 2010.
Book review of 'Field Techniques for Sea Ice Research' by H Eicken et al. Polar Biology. 33, 1161-1162.

Thomas, D.N. 2005
Antarctic Marine Protists. FJ Scott and H Marchant. Polar Record. 41, 366-367.

Thomas, D.N. 2005
Life in Ancient Ice. JD Castello and SO Rogers. The Quarterly Review of Biology. 80(4),495.

Thomas, D.N. 2004
The Snowflake. K Libbrecht and P Rasmussen. BBC Wildlife Magazine vol. (December).

Thomas, D.N. 2001
Plant Life. R Ennos and E Sheffield. BBC Wildlife Magazine vol. 19 (February).

Thomas, D.N. 1999
The Biology of Polar Habitats. GE Fogg. BBC Wildlife Magazine vol. 17 (April).

Thomas, D.N. 1996
Algae: An Introduction to Phycology. C van den Hoek, DG Mann and HM Jahns. BBC Wildlife Magazine vol. 14 (September).

Thomas, D.N. 1996
Freshwater algae: Their microscopic world explored. H Canter- Lund and JWG Lund. BBC Wildlife Magazine vol. 14 (April).

Book Chapters

DN Thomas, H Kaartokallio, L Tedesco, J Piiperinen, M Majaneva, J-M Rintala, E Eronen, H Kuosa, J Blomster and MA Granskog. 2014.
Chapter 10 - Sea ice. In: Biological Oceanography of the Baltic Sea (eds. P Snoeijs, H Schubert and T Radziejewska). Springer. ISBN 978-94-007-0667-5
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DN Thomas. 2012. Chapter 4 - Sea ice. In: Life at Extremes: Environments, Organisms and Strategies for Survival (ed. EM Bell). CABI Publishing. Wallingford, U.K. pp 62-80.

P Convey, DKA Barnes, H Griffiths, S Grant, K Linse and DN Thomas. 2012.
Chapter 15 - Biogeography and regional classifications of Antarctica. In: Antarctic Ecosystems: An Extreme Environment in a Changing World (eds. AD Rogers, NM Johnston, EJ Murphy and A Clarke). Blackwell Publishing. pp 461-480.

DN Thomas. 2011.
Sea ice. In: Life at Extremes: Environments, Organisms and Strategies for Survival (ed. EM Bell). CABI Publishing. Wallingford, U.K. ISBN-13: 9781845938147
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DN Thomas. 2011.
The Biogeochemistry of sea ice. In: Encyclopedia of Snow, Ice and Glaciers (Eds. U Haritashya, P Singh and VP Singh). Springer, Dordrecht Heidelberg. pp. 98-102.

DN Thomas. 2011.
The Nature of Ice. In: Arctic Tipping Points (eds. P Wassmann and CM Duarte). BBVA Foundation, Bilbao, Spain. 50-51.

P Convey, DKA Barnes, H Griffiths, S Grant, K Linse and DN Thomas. 2011.
Biogeography and regional classifications of Antarctica. In: Antarctic Ecosystems: An Extreme Environment in a Changing World (eds. AD Rogers, NM Johnston, EJ Murphy and A Clarke). Wiley. DOI: 10.1002/9781444347241
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Thomas, DN, Papadimitriou, S and Michel, C. 2010.
The Biogeochemistry of sea ice. In: Sea ice - 2nd Edition (Eds. DN Thomas and GS Dieckmann) Wiley-Blackwell Publishing.

GS Dieckmann and DN Thomas. 2009.
Sea ice: an icy crucible. In: Biological studies in Polar Oceans: Exploration of life in icy waters (Eds. I and G Hempel). Wirtschaftsverlag. Nordwest Wilhelmshaven, Germany.

Mock, T and Thomas, DN. 2007.
Microalgae in Polar regions: Linking functional genomics and physiology with environmental conditions. In: Psychrophiles: From Biodiversity to Biotechnology (Eds. R Margesin, F Schinner, J-C Marx and C Gerday). Springer Verlag, Berlin. pp. 285-312.

Kaiser, M.J., Hall, S.J. and Thomas, D.N.. 2005
Habitat modification. In: The Sea, volume 13 (Eds. AL Robinson and KH Brink). Harvard University Press. pp. 927-970.
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Lara, R.J., U Hubberten, DN Thomas and G Kattner. 1996
Humic substance research in polar waters. In: Humic substances and organic matter in soil and water environments: Characterisation. Transformation and interactions. (Eds. C Clapp, MHB Hayes, N Senesi and S Griffith). pp. 241-248.

Additional Publications

Papadimitriou S, Kennedy H, Kennedy P, Thomas DN (2014)
Seawater freezing and hydrated sea salt formation: Ikaite in the sea ice of polar oceans. In: Diamond Light Source Annual Review 2013/14, Publisher: Diamond Light Source, pp.42-43

DN Thomas. 2011. Studying the natural history of pack ice in the Weddell Sea. In: Nimrod, The Journal of the Ernest Shackleton Autumn School, 5, 20-26.

J.M. Webb, L. Le Vay, D.N. Thomas and M. Rigby 2009
Integrated hydroponics: Growing Samphire in wastewater from a marine fish farm (Article) Finfish News 7, 9-12.

Thomas, D.N. and Mock, T. (2005)
Life in frozen veins - Coping with the cold. The Biochemist, 27, 12-16.

Thomas, D.N. 2004
From Plankton to Trees. Form Magazine vol 1 (December), 17-21

Thomas, D.N. 2004
La banquisa: 'Pack Ice'. Omnis cellula vol 6, 9-15 (translated into Catalan)

Thomas, D.N. 2003
Tough diamonds. BBC Wildlife Magazine vol.21 (May).

Thomas, D.N. 2002
Strangers in a Strange Land. Kew Magazine Vol. 37. 24-27

Thomas, D.N., Dieckmann, G. and Mock, T. 2002
Lebensraum Meereis. Biologen Heute, vdbiol.

Dieckmann, G., Mock, T. and Thomas, D.N. 2002
Meereis. Spektrum der Wissenschaft Dossier 3/2002.

Thomas, D.N. 2001
Ice Station Hamburg. NERC News. Autumn 2001

Thomas, D.N. 2001
Marine craft finds bilge pump useful. The Times Higher Education Supplement. 07.09.01

Thomas, D.N. 2001
Wonder Weeds. BBC Wildlife Magazine vol. 19 (April).

Thomas, D.N. 2000
How we can tell mummy ate fish. The Times Higher Education Supplement 11.08.00.

Thomas, D.N. 2000
Salt Removal - Last Word. New Scientist 21.10.00.

Thomas, D.N. 2000
Saving fragile forests. The Times Higher Education Supplement 18.02.00.

Thomas, D.N. 2000
The art of nature. BBC Wildlife Magazine vol.18 (July).

Thomas, D.N. 1999
Don's Diary. The Times Higher Education Supplement 21.05.99.

Thomas, D.N. 1999
Don's Diary. The Times Higher Education Supplement 27.08.99.

Thomas, D.N. 1999
Hot questions of shelf life. The Times Higher Education Supplement 17.09.99.

Thomas, D.N. 1999
Life as a BA media fellow. NERC News. Winter 1999.

Thomas, D.N. 1999
Living Gems. BBC Wildlife Magazine vol.17 (February).

Thomas, D.N. 1999
Living in sea ice. NERC News. Summer 1999.

Thomas, D.N. 1999
No quick fix for excesses. BBC Wildlife Magazine vol.17 (May).

Thomas, D.N. 1999
Science loses out to Brit. hype. The Times Higher Education Supplement 03.09.99.

Thomas, D.N. 1999
Search for the secrets of Arctic ice floes. The Times Higher Education Supplement 10.03.99.

Thomas, D.N. 1998
Don's Diary. The Times Higher Education Supplement 13.03.98.

Thomas, D.N. 1998
The Big Chill. Daily Post. 19.03.98.

Russell, G. and Thomas, D.N. 1988
The Baltic Sea: A cradle of plant evolution? Plants Today 1. 77-82.

Thomas, D.N. 1996
Nature's ice- show. BBC Wildlife Magazine vol.14 (December).

Thomas, D.N. and Dieckmann, G.S. 1994
Life in a frozen lattice. New Scientist 142.

Thomas, D.N. 1993
Splat Count. Letter to New Scientist. 4.12.93

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