Professor Lewis LeVay

Photograph of Prof Lewis LeVay


Director of the Centre for Applied Marine Sciences (CAMS), with a background in aquaculture, fisheries and ecology across a wide range of temperate and tropical environments. After a B.Sc. in Biology, Sussex University and MSc in Marine Biology (Bangor), I worked for several years on shrimp and fish nutrition and feed development, in collaboration with several industrial partners. The results from this research contributed to my PhD from Bangor University, which was followed by several years working in industry and as a consultant. I returned to full time research working in SE Asia on CAMS projects developing techniques for mangrove-integrated aquaculture and fisheries stock enhancement, which led to an academic position in the School of Ocean Sciences and then to my current dual role as Director of CAMS and Professor in the School, being awarded a personal chair in October 2014.

Research Interests

Recent aquaculture research includes participation in the long-standing Atlantic Area SEAFARE consortium, with projects on integration of aquaculture in coastal wetlands and wastewater remediation in constructed wetlands, with further projects on development of offshore aquaculture for seaweeds and shellfish. I have long-term ongoing research programmes on hatchery technologies, feeds and nutrition, and aquaculture for tropical species including penaeid shrimp mud crabs Scylla spp. Working on development of stock enhancement strategies has led to more ecological research in tropical and subtropical systems, principally on mangroves and seagrasses in SE Asia, East Africa and the Arabian Gulf. Current fisheries work is focused on shellfish fisheries in Welsh waters through the Sustainable Fisheries for Wales project (EFF-funded) and a new project with Qatar University on ecosystem approaches to management of demersal fisheries in the Arabian Gulf. As director of CAMS, I am also closely involved in SEACAMS, a major Welsh convergence initiative that links marine science expertise with SMEs, commercial and institutional partners in marine sector industries such as renewable energy, infrastructure development, engineering and tourism.

Research Groups

Research Themes

Research Grants Awarded

Transnational approaches to resolving biological bottlenecks in macroalgal biofuel production (SuBBSea) BBSRC-DBT(India) 2015-2017

Identification of essential fish habitat to achieve an ecosystem-based approach to fisheries management (with Qatar University and Qatar Ministry of Environment, Department of Fisheries, funded by Qatar National Research Foundation, National Priorities Research Programme) (2013-2016)

Ecological processes underlying ecosystem function in arid mangroves (with Qatar, funded by Qatar National Research Foundation, National Priorities Research Programme) (2015-2018)

Sustainable aquaculture for the Atlantic Region of Europe (SEAFARE) (2009-2013)

Sustainable fisheries for Wales (2012-2015)

Novel ingredients from seaweeds (NISE) (2012-2013)

Nitrogen and carbon cycling in shallow water marine habitats (with Qatar University, funded by Qatar National Research Foundation, National Priorities Research Programme) (2009-2013)

Identification of essential fish habitat to achieve an ecosystem-based approach to fisheries management (with Qatar University and Qatar Ministry of Environment, Department of Fisheries, funded by Qatar National Research Foundation, National Priorities Research Programme) (2013-2016)

Swahili Seas, NERC-DFID Ecosystem services for poverty alleviation (2011-2013)

Low carbon research initiative (LCRI)


Peer Reviewed Journal Papers

Galley TH, Beaumont AR, Le Vay L, King, J., 2017.
Influence of exogenous chemicals on larval development and survival of the king scallop Pecten maximus (L.). Aquaculture, 474, pp.48–56.

Mark E. M. Walton, Jamie Hayes, Mohsin Al-Ansi, Mohamed Abdallah, Ibrahim Al Maslamani, Mohammed Al-Mohannadi, Ismail Al-Shaikh, Tim D'Urban Jackson, Claire Szostek, Jack Egerton, Michel J. Kaiser, Lewis Le Vay (2017).
Towards Spatial Management of Fisheries in the Arabian Gulf: Benthic diversity, habitat and fish distributions from Qatari waters. ICES Journal of Marine Science.

González-Ortegón E., L Gimánez, J Blasco and L Le Vay (2015)
Effects of food limitation and pharmaceutical compounds on the larval development and morphology of Palaemon serratus. Science of the Total Environment 503, 171-178

Quintã R, R Santos, DN Thomas and L Le Vay. 2015.
Growth and nitrogen uptake by Salicornia europaea and Aster tripolium in nutrient conditions typical of aquaculture wastewater. Chemosphere,120, 414-421

Quintã, R, P Hill, D. Jones, R. Santos, DN Thomas and L Le Vay. 2015.
Uptake of an amino acid (alanine) and its peptide (trialanine) by the saltmarsh halophytes Salicornia europaea and Aster tripolium and its potential role in ecosystem N cycling and marine aquaculture wastewater treatment. Environmental Engineering, 75, 145-154.

González-Ortegón E., Walton M.E.M., Moghaddam B., Vilas C., Prieto A., Kennedy H.A., Cañavate J.P and Le Vay L.  (2015)
Flow regime in a restored wetland determines trophic links and species composition in the aquatic macroinvertebrate community. Science of The Total Environment 503, 241-250

Walton M.E.M, Vilas C, Cañavate JP, González-Ortegón E., Prieto Ab, van Bergeijk S.A, Green A.J, Librero M,. Mazuelos M. and Le Vay L (2015).
A model for the future: Ecosystem services provided by the aquaculture activities of Veta la Palma, Southern Spain. Aquaculture 448: 382-390

Walton M.E.M., Vilas C. ,Coccia C., Green A.J., Cañavate J.P.,  Prieto A., van Bergeijk S.A., Medialdea J.M.,  Kennedy H., King, J. and Le Vay L. (2015)
The effect of water management on extensive aquaculture food webs in the reconstructed wetlands of the Doñana Natural Park, Southern Spain. Aquaculture 448: 451- 463

Walton M.E.M., I. Al-Maslamani, M.W. Skov, I. Al-Shaikh, I.S. Al-Ansari, H.A. Kennedy, L. Le Vay (2014)
Outwelling from arid mangrove systems is sustained by inwelling of seagrass productivity. Marine Ecology Progress Series. DOI: 10.3354/meps10827.

Jones A.C., Mead A., Kaiser M.J., Austen M.C.V., Adrian A.W., Auchterlonie N.A., Black K.D., Blow L.R., Bury C., Brown J.H., Burnell G.A., Connolly E., Dingwall A., Derrick S., Eno N.C., Gautier D.J.H., Green K.A., Gubbins M., Hart P.R., Holmyard J.M., Immink A.J., Jarrad D.L., Katoh E., Langley J.C.R., Lee D. O'C., LeVay L., Leftwich C.P., Mitchell M., Moore A.P., Murray A.G., McLaren E.M.R., Norbury H., Parker D., Parry S.O., Purchase D., Rahman A., Sanver F., Siggs M., Simpson S.D., Slaski R.J., Smith K., Syvret M. LeQ., Tibbott C., Thomas P.C., Turnbull J., Whiteley R., Whittles M., Wilcockson M.J., Wilson J., Dicks L.V., and Sutherland W.J. (2014)
Prioritisation of knowledge-needs for sustainable aquaculture: a national and global perspective. Fish and Fisheries. DOI: 10.1111/faf.12086

Al-Maslamani, M.E.M. Walton, H.A. Kennedy, M. Al-Mohannadi, L. Le Vay (2013)
Are mangroves in arid environments isolated systems? Life-history and evidence of dietary contribution from inwelling in a mangrove-resident shrimp species. Estuarine Coastal Shelf Science 124: 56-63

E González-Ortegón, J Blasco, L Le Vay, L Giménez (2013)
A multiple stressor approach to study the toxicity and sub-lethal effects of pharmaceutical compounds on the larval development of a marine invertebrate. Journal of Hazardous Materials 263P (2013) 233-238

JM Webb, R Quintã, S Papadimitriou, L Norman, M Rigby, DN Thomas, L Le Vay (2013)
The effect of halophyte planting density on the efficiency of constructed wetlands for the treatment of wastewater from marine aquaculture Ecological Engineering 61, 145-15

Al-Maslamani I., Kennedy H., Walton M.E.M. and Le Vay L. (2012)
Sources of benthic primary production supporting food webs in an arid coastal embayment. Marine Biology 159: 1753-1762

JM Webb, R Quintã, S Papadimitrou, L Norman, M Rigby, DN Thomas and L LeVay. 2012.
Halophyte filter beds for treatment of saline wastewater from aquaculture. Water Research, 46, 5102-5114.

Lebata J.H., Walton M.E., Biñas J.B., Primavera J.H. and Le Vay L. (2012)
Identifying mangrove areas for fisheries enhancement; population assessment in a patchy habitat. Aquatic Conservation 22: 652-664

Galley T., Green B., Watkins L., Le Vay L. (2011)
Optimising hatchery production of the European shore crab Carcinus maenas. Aquaculture International 19: 381-394

Gamboa-Delgado, J., Le Vay, L., Manchado, M., Ponce, M., Fernandez-Diaz, C., Zerolo, R.,Cañavate, J.P. (2011)
Effect of larval feeding regime on trypsinogen gene expression, proteolytic enzyme activity and dietary carbon and nitrogen assimilation in Senegalese sole (Solea senegalensis) larvae. Comparative Biochemistry and Physiology Part B 158, 251-258

Hother-Parkes L., Quinitio E.T., Le Vay L. (2011)
Phenotypic differences between hatchery-reared and wild mud crabs, Scylla serrata, and the effects of conditioning. Aquaculture International 19: 3610-380

Le Vay, L. and Gamboa-Delgado, J. (2011)
Naturally-occurring stable isotopes as direct measures of larval feeding efficiency, nutrient incorporation and turnover. Aquaculture 315, 95-103

Primavera J.H., Binas J.B., Samonte-Tan G.B.P., Lebata J., Alava V.R., Walton M., LeVay L. (2010)
Mud crab pen culture: replacement of fish feed requirement and impacts on mangrove community structure. Aquaculture Research 41 : 1211-1220

Al-Maslamani, I.; Le Vay, L., Kennedy, H. (2009)
Feeding on intertidal microbial mats by postlarval tiger shrimp, Penaeus semisulcatus De Haan. Marine Biology 156: 2001-9

Gamboa Delgado J. and Le Vay L (2009)
Artemia replacement in co-feeding regimes for mysis and postlarval stages of Litopenaeus vannamei: Nutritional contribution of inert diets to tissue growth as indicated by natural carbon stable isotopes. Aquaculture 297: 128-135

Gamboa-Delgado J. and Le Vay L. (2009)
Natural stable isotopes as indicators of the relative contribution of soy protein and fish meal to tissue growth in Pacific white shrimp (Litopenaeus vannamei) fed compound diets. Aquaculture 291: 115-123

Lebata JHL, Le Vay L., Walton M.E., Biñas J.B, Quinitio E.T., Rodriguez E.M., Primavera J.H. (2009)
Evaluation of hatchery-based enhancement of the mud crab, Scylla spp., fisheries: comparison of species and release strategies. Mar. Fresh Res. 60: 58-69

Gamboa-Delgado, J., Ca, Cañavate, J. P., Zerolo, R. and Le Vay, L. 2008.
Natural carbon stable isotope ratios as indicators of the relative contribution of live and inert diets to growth in larval Senegalese sole (Solea senegalensis). Aquaculture 280, 1-4, Pages 190-197.
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Le Vay L., Lebata J.H.L., Walton M., Primavera J.H., Quinitio E.T., Lavilla-Pitigo C., Parado-Estepa F., Rodriguez E., Vu Noc Ut, Truong Trong Nghia, Sorgeloos P. and Wille M. (2008)
Approaches to enhancement in mangrove-associated crab fisheries. Reviews in Fisheries Science 16:1, 72 - 80

Lebata J.H., Le Vay L. , Primavera, J.H. , Walton M. and Biñas J.
Enhancement of fisheries for mud crabs Scylla spp. in the mangroves of Naisud and Bugtong, Ibajay, Aklan, Philippines Baseline assessment of species abundance. Bulletin of Marine Science 80: 891-904

Ut V.N., Le Vay L., Nghia TT. (2008)
Development of nursery culture techniques for the mud crab Scylla paramamosain. Aquaculture Research 38: 1563-1568

Al-Maslamani I, Le Vay, L, Kennedy, H and Jones D.A.(2007)
Feeding ecology of the grooved tiger shrimp Penaeus semisulcatus De Haan in the coastal waters of Qatar, Arabian Gulf. Marine Biology 150: 627-637

Le Vay L., Carvalho G., Quinitio E.T., Lebata J.H., Ut V.N. and Fushimi H. (2007)
Quality of hatchery-reared juveniles for marine fisheries stock enhancement. Aquaculture 268: 169-180

Le Vay L., Ut V.N. and Walton M. (2007)
Population ecology of the mud crab Scylla paramamosain (Estampador) in an estuarine mangrove system; a mark-recapture study. Marine Biology151: 1127-1135.

Ut V.N., Le Vay L., Nghia T.T., Walton M. (2007)
Comparative performance of hatchery- reared and wild Scylla paramamosain in pond culture. Aquaculture Research 38: 1593-1599

Walton M., Le Vay L, Lebata J.H., Binas J., Tapper J., Primavera J.H. (2007)
Effectiveness of mangrove replanting in the restoration of fisheries: abundance and distribution of mud crabs, Scylla olivacea in intact, degraded and rehabilitated mangroves. Biological Conservation 138: 180-188

Walton M.E.M., Samonte G., Primavera JH, Edwards-Jones G., Le Vay L. (2006)
Are mangroves worth replanting? The direct economic benefits of a community-based reforestation project. Environmental Conservation 33: 335-343

Walton, M.E.M., Le Vay, L., Lebata, J.H., Binas, J. and Primavera, J.H. (2006)
Seasonal abundance, distribution and recruitment of mud crabs (Scylla spp.) in replanted mangroves. Estuarine, Coastal and Shelf Science, 66, 493- 500.

Walton, M.E.M., LeVay, L., Truong, L.M. and Ut, V.N. (2006)
Significance of mangrove-mudflat boundaries as nursery grounds for the mud crab, Scylla paramamosian. Marine Biology, DOI: 10.1007/s00227-006-0267-7.

Al-Jamali, F; Bishop, JM; Osment, J; Jones, DA; Le Vay, L (2005)
A review of the impacts of aquaculture and artificial waterways upon coastal ecosystems in the Gulf (Arabian/Persian) including a case study demonstrating how future management may resolve these impacts. Aquatic Ecosystem Health and Management. 8: 81-94.

Puello-Cruz A.C., Sangha R.S., Jones D.A. and Le Vay L. (2002)
Trypsin enzyme activity during larval development of Litopenaeus vannamei (Boone) fed on live feeds. Aquaculture Research. 33: 333-338

Le Vay, L. (2001)
Ecology and management of mud crabs, Scylla spp. Asian Fisheries Science, 14, 101–111.

Le Vay, L., Jones, D.A., Puello-Cruz, A.C., Sangha, R.S. and Ngamphongsai, C. (2001)
Digestion in relation to feeding strategies exhibited by crustacean larvae. Comparative Biochemistry and Physiology, Part A, 128, 623–630.

Le Vay, L., Vu Ngoc Ut and Jones, D.A. (2001)
Seasonal abundance and recruitment in an estuarine population of mud crabs, Scylla paramamosian, in the Mekong Delta, Vietnam. Hydrobiologia, 449, 231–239.

Ut V.N., Le Vay L., Nghia T.T., Hanh T.T.H. and Caldwell B.S. (2001)
Effect of diet and substrate in the nursery phase culture of mud crab (Scylla paramamosain) production. Europ. Aquaculture Soc. Spec. Publ. 30: 610 -613

Jones D.A., Kumlu M., Le Vay L. and Fletcher D.J. (1997)
The digestive physiology of herbivorous, omnivorous and carnivorous crustacean larvae. Aquaculture 155: 285 - 295

Kamarudin M.S., Jones D.A., Le Vay L. and A. Z. Abidan (1994)
Ontogenetic changes in digestive enzyme activity during larval development of Macrobrachium rosenberbgii. Aquaculture123: 323-333.

Rodríguez, A., Le Vay, L., Mourente, G. and Jones D.A. (1994)
Biochemical composition and digestive enzyme activity in larvae and postlarvae of Penaeus japonicus (Bate), during herbivorous and carnivorous feeding. Mar. Biol. 118: 45-53

Jones, D.A., Kamarudin, M.S. and Le Vay, L. (1993).
The potential for replacement of live feeds in larval culture. Journal World Aquaculture Society 24: 199-210

Le Vay, L., Rodríguez, A., Kamarudin, M.S. and Jones, D.A. (1993)
Influence of live and artificial diets on tissue composition and digestive enzyme activity in Penaeus japonicus larvae. Aquaculture 118: 287-297

Additional Publications

Walton MEM, Vilas C, Cañavate JP, Prieto A, van Bergeijk SA, Medialdea JM, Librero M, Mazuelos N, King JW, Lee D. O'C and Le Vay L (2015) Policy Guidance for Sustainable Wetlands Aquaculture. SEAFARE Policy report. DOI: 10.13140/RG.2.1.4579.7203

J.M. Webb, L. Le Vay, D.N. Thomas and M. Rigby 2009
Integrated hydroponics: Growing Samphire in wastewater from a marine fish farm (Article) Finfish News 7, 9-12.

Lee, D, King JW, Levay, L, McCartney, I and Beaumont, A (2008)
Atlantic Arc Aquaculture Group Final Report. Centre for Applied Marine Sciences. Report 2008-08.

PDF copies of some of these publications are available (as indicated by hyperlinks in the list) for non-commercial research and educational purposes only. You may not use the material for commercial purposes or make multiple copies without requesting permission from the copyright holder (usually the book or journal publisher).